==== Secondary Structure Definition by the program DSSP, updated CMBI version by ElmK / April 1,2000 ==== DATE=20-JUL-2011 . REFERENCE W. KABSCH AND C.SANDER, BIOPOLYMERS 22 (1983) 2577-2637 . HEADER HYDROLASE 21-JUL-05 2BX2 . COMPND 2 MOLECULE: RIBONUCLEASE E; . SOURCE 2 ORGANISM_SCIENTIFIC: ESCHERICHIA COLI; . AUTHOR M.J.MARCAIDA,A.J.CALLAGHAN,W.G.SCOTT,B.F.LUISI . 500 4 0 0 0 TOTAL NUMBER OF RESIDUES, NUMBER OF CHAINS, NUMBER OF SS-BRIDGES(TOTAL,INTRACHAIN,INTERCHAIN) . 26753.0 ACCESSIBLE SURFACE OF PROTEIN (ANGSTROM**2) . 341 68.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(J) , SAME NUMBER PER 100 RESIDUES . 28 5.6 TOTAL NUMBER OF HYDROGEN BONDS IN PARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 83 16.6 TOTAL NUMBER OF HYDROGEN BONDS IN ANTIPARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 1 0.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-5), SAME NUMBER PER 100 RESIDUES . 2 0.4 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-4), SAME NUMBER PER 100 RESIDUES . 3 0.6 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-3), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-2), SAME NUMBER PER 100 RESIDUES . 1 0.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-1), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+0), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+1), SAME NUMBER PER 100 RESIDUES . 40 8.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+2), SAME NUMBER PER 100 RESIDUES . 56 11.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+3), SAME NUMBER PER 100 RESIDUES . 121 24.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+4), SAME NUMBER PER 100 RESIDUES . 5 1.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+5), SAME NUMBER PER 100 RESIDUES . 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 *** HISTOGRAMS OF *** . 0 0 0 2 3 1 1 0 1 1 0 0 1 1 0 0 1 0 0 0 0 1 1 0 0 0 0 0 0 0 RESIDUES PER ALPHA HELIX . 1 1 1 1 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PARALLEL BRIDGES PER LADDER . 0 1 3 4 1 4 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 ANTIPARALLEL BRIDGES PER LADDER . 0 1 1 1 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 LADDERS PER SHEET . # RESIDUE AA STRUCTURE BP1 BP2 ACC N-H-->O O-->H-N N-H-->O O-->H-N TCO KAPPA ALPHA PHI PSI X-CA Y-CA Z-CA 1 -6 L A 0 0 131 0, 0.0 0, 0.0 0, 0.0 0, 0.0 0.000 360.0 360.0 360.0 45.9 93.0 -79.8 -1.9 2 -5 L H + 0 0 96 1,-0.0 2,-0.4 4,-0.0 0, 0.0 -0.144 360.0 178.9 -51.6 129.6 91.0 -76.7 -3.0 3 -4 L I > - 0 0 100 0, 0.0 3,-1.6 0, 0.0 4,-0.4 -0.994 34.3-115.0-129.6 128.1 93.0 -74.0 -5.0 4 -3 L E T 3 S+ 0 0 113 -2,-0.4 23,-0.0 1,-0.2 -2,-0.0 -0.427 101.9 29.8 -66.4 126.4 91.3 -70.9 -6.3 5 -2 L G T 3 S+ 0 0 60 2,-0.3 -1,-0.2 -2,-0.2 4,-0.1 -0.183 90.2 97.5 120.0 -38.1 91.4 -71.1 -10.1 6 -1 L R S < S+ 0 0 135 -3,-1.6 2,-0.2 2,-0.1 -2,-0.1 0.940 86.8 29.7 -49.7 -64.6 91.3 -74.8 -10.7 7 0 L H S S- 0 0 101 -4,-0.4 -2,-0.3 1,-0.1 2,-0.1 -0.509 96.8 -86.6 -95.2 163.5 87.6 -75.3 -11.3 8 1 L M - 0 0 62 -2,-0.2 20,-2.8 19,-0.1 2,-0.5 -0.427 34.4-135.6 -60.3 137.0 84.9 -73.0 -12.8 9 2 L K E -Ab 27 222A 21 212,-0.6 214,-2.0 18,-0.3 215,-0.7 -0.836 33.3-174.3 -84.8 133.9 83.2 -70.6 -10.4 10 3 L R E -Ab 26 224A 46 16,-3.7 16,-2.9 -2,-0.5 2,-0.7 -0.945 29.2-134.5-130.8 156.5 79.5 -70.7 -11.2 11 4 L M E -Ab 25 225A 1 213,-2.6 215,-2.0 -2,-0.3 2,-0.6 -0.925 25.4-164.3-104.6 113.2 76.4 -68.9 -10.1 12 5 L L E -Ab 24 226A 0 12,-3.3 12,-3.4 -2,-0.7 2,-0.6 -0.871 4.0-168.7-100.8 123.5 73.6 -71.4 -9.5 13 6 L I E -Ab 23 227A 0 213,-2.7 215,-1.6 -2,-0.6 2,-0.8 -0.929 1.9-166.6-113.4 111.0 70.1 -70.1 -9.3 14 7 L N E +A 22 0A 15 8,-2.4 8,-2.2 -2,-0.6 3,-0.1 -0.845 23.4 153.1 -98.8 109.5 67.5 -72.6 -8.0 15 8 L A + 0 0 13 -2,-0.8 217,-0.1 6,-0.2 -1,-0.1 -0.066 34.1 107.4-131.1 33.4 64.0 -71.2 -8.7 16 9 L T S S+ 0 0 68 4,-0.1 2,-0.5 1,-0.1 -1,-0.1 0.594 71.6 68.2 -83.1 -9.4 61.7 -74.2 -9.0 17 10 L Q > - 0 0 104 3,-0.3 3,-1.6 -3,-0.1 -1,-0.1 -0.938 69.9-151.5-117.8 115.5 60.1 -73.6 -5.6 18 11 L Q T 3 S+ 0 0 141 -2,-0.5 3,-0.2 1,-0.3 -1,-0.1 0.868 91.9 55.7 -53.1 -37.8 57.8 -70.5 -5.2 19 12 L E T 3 S+ 0 0 107 1,-0.2 -1,-0.3 19,-0.1 2,-0.2 0.532 113.2 30.3 -85.2 1.0 58.5 -70.1 -1.5 20 13 L E < - 0 0 84 -3,-1.6 2,-0.5 18,-0.1 -3,-0.3 -0.741 62.3-164.0-160.4 103.6 62.3 -69.9 -1.5 21 14 L L E - C 0 37A 41 16,-4.6 16,-3.0 -2,-0.2 2,-0.5 -0.788 20.2-161.1 -87.6 120.2 64.4 -68.5 -4.3 22 15 L R E -AC 14 36A 24 -8,-2.2 -8,-2.4 -2,-0.5 2,-0.5 -0.884 16.1-173.2-117.4 132.8 68.1 -69.5 -3.9 23 16 L V E -AC 13 35A 8 12,-3.9 12,-1.8 -2,-0.5 2,-0.4 -0.958 11.8-178.1-122.0 120.9 71.3 -68.1 -5.3 24 17 L A E -AC 12 34A 0 -12,-3.4 -12,-3.3 -2,-0.5 2,-0.7 -0.933 22.5-149.6-120.9 134.5 74.6 -69.9 -4.8 25 18 L L E -AC 11 33A 10 8,-2.4 7,-2.7 -2,-0.4 8,-1.4 -0.911 28.9-175.3 -93.1 121.7 78.1 -69.1 -5.8 26 19 L V E -AC 10 31A 1 -16,-2.9 -16,-3.7 -2,-0.7 2,-0.6 -0.930 22.5-157.0-119.0 142.0 79.9 -72.4 -6.3 27 20 L D E > S-AC 9 30A 5 3,-2.8 3,-3.0 -2,-0.4 2,-0.5 -0.948 77.5 -46.1-116.8 110.0 83.5 -73.1 -7.1 28 21 L G T 3 S- 0 0 9 -20,-2.8 -18,-0.1 -2,-0.6 -2,-0.0 -0.585 124.7 -32.0 60.3-116.3 83.7 -76.5 -8.7 29 22 L Q T 3 S+ 0 0 104 -2,-0.5 2,-0.8 -3,-0.1 -1,-0.3 0.097 116.8 111.2-111.1 19.6 81.4 -78.3 -6.2 30 23 L R E < -C 27 0A 75 -3,-3.0 -3,-2.8 240,-0.0 2,-0.3 -0.838 62.4-142.6-101.1 110.1 82.5 -76.1 -3.3 31 24 L L E +C 26 0A 0 -2,-0.8 -5,-0.2 -5,-0.2 3,-0.1 -0.533 30.0 163.7 -67.7 127.0 79.7 -73.8 -2.1 32 25 L Y E + 0 0 22 -7,-2.7 2,-0.3 1,-0.4 -6,-0.2 0.473 61.2 20.7-125.1 -5.9 81.2 -70.4 -1.1 33 26 L D E -C 25 0A 24 -8,-1.4 -8,-2.4 295,-0.1 -1,-0.4 -0.956 58.8-171.8-160.3 149.6 78.2 -68.2 -0.9 34 27 L L E -C 24 0A 10 -2,-0.3 2,-0.4 -10,-0.2 -10,-0.2 -0.984 5.9-177.0-140.9 144.0 74.5 -68.4 -0.6 35 28 L D E -C 23 0A 49 -12,-1.8 -12,-3.9 -2,-0.4 2,-0.4 -0.988 7.1-171.8-143.7 138.0 71.5 -66.2 -0.8 36 29 L I E -C 22 0A 48 -2,-0.4 2,-0.4 -14,-0.2 -14,-0.2 -0.991 3.2-178.4-135.6 128.1 67.9 -67.1 -0.2 37 30 L E E -C 21 0A 27 -16,-3.0 -16,-4.6 -2,-0.4 5,-0.1 -0.980 1.1-176.0-130.0 141.3 64.8 -65.1 -0.8 38 31 L S - 0 0 42 -2,-0.4 3,-0.4 3,-0.3 -19,-0.1 -0.991 38.8 -88.2-135.5 144.8 61.1 -65.8 -0.2 39 32 L P S S+ 0 0 97 0, 0.0 2,-1.2 0, 0.0 -21,-0.0 -0.143 109.7 14.3 -45.8 137.0 57.9 -63.9 -0.9 40 33 L G S S+ 0 0 81 1,-0.1 2,-1.2 0, 0.0 0, 0.0 -0.455 111.6 73.8 95.3 -62.0 56.9 -61.5 1.9 41 34 L H S S+ 0 0 87 -2,-1.2 -3,-0.3 -3,-0.4 -1,-0.1 -0.154 72.7 157.3 -78.1 42.7 60.2 -61.5 3.8 42 35 L E - 0 0 62 -2,-1.2 2,-0.1 1,-0.1 166,-0.1 -0.302 48.4 -86.4 -66.9 154.9 61.7 -59.3 1.0 43 36 L Q - 0 0 37 1,-0.1 163,-0.2 164,-0.1 -1,-0.1 -0.318 34.6-177.9 -68.8 142.0 64.8 -57.1 1.7 44 37 L K > + 0 0 28 161,-0.1 3,-1.5 -2,-0.1 2,-0.3 0.502 46.5 112.4-116.0 -10.5 64.4 -53.5 3.1 45 38 L K T 3 S+ 0 0 47 1,-0.3 61,-0.1 340,-0.1 3,-0.1 -0.498 83.5 18.8 -66.4 126.8 68.1 -52.4 3.2 46 39 L A T 3 S+ 0 0 22 59,-1.2 -1,-0.3 1,-0.4 2,-0.2 0.606 88.1 137.9 83.4 12.4 68.8 -49.7 0.7 47 40 L N < - 0 0 4 -3,-1.5 58,-1.8 158,-0.1 2,-0.4 -0.516 45.3-138.8 -67.3 155.1 65.2 -48.7 0.3 48 41 L I E +EF 104 204B 3 156,-2.2 155,-4.1 56,-0.2 156,-1.8 -0.979 24.1 175.5-125.6 139.1 64.6 -44.9 0.2 49 42 L Y E -EF 103 202B 27 54,-3.1 54,-2.4 -2,-0.4 2,-0.7 -0.817 37.9-117.9-132.2 168.6 61.8 -43.0 1.9 50 43 L K E -EF 102 201B 7 151,-3.0 151,-1.2 17,-0.3 2,-0.3 -0.937 48.1-175.6-105.0 103.5 60.4 -39.5 2.6 51 44 L G E -E 101 0B 0 50,-2.2 50,-0.9 -2,-0.7 2,-0.4 -0.763 22.3-141.2-108.1 148.5 60.6 -39.4 6.4 52 45 L K E -EG 100 66B 25 14,-1.7 14,-2.7 -2,-0.3 2,-0.4 -0.896 31.2-109.9-105.1 134.4 59.4 -36.9 9.0 53 46 L I E - G 0 65B 10 46,-1.6 45,-2.7 -2,-0.4 12,-0.3 -0.474 34.7-176.8 -65.4 115.2 61.5 -36.2 12.1 54 47 L T E - 0 0 38 10,-2.8 2,-0.3 -2,-0.4 -1,-0.2 0.983 60.2 -9.4 -74.8 -75.7 59.7 -37.7 15.1 55 48 L R E - 0 0 110 9,-0.2 9,-2.3 42,-0.1 -1,-0.3 -0.971 64.1-135.0-134.0 145.3 61.8 -36.8 18.2 56 49 L I E - G 0 63B 43 -2,-0.3 7,-0.2 7,-0.2 3,-0.1 -0.606 11.3-165.5-101.5 160.3 65.2 -35.2 18.7 57 50 L E E >> + G 0 62B 52 5,-1.2 5,-1.1 -2,-0.2 4,-1.0 -0.808 20.8 174.4-147.7 100.5 68.1 -36.2 21.0 58 51 L P T 45S+ 0 0 83 0, 0.0 3,-0.2 0, 0.0 -1,-0.1 0.847 87.1 69.7 -68.2 -28.7 71.1 -34.1 21.8 59 52 L S T 45S+ 0 0 97 1,-0.3 -2,-0.0 -3,-0.1 -3,-0.0 0.874 113.4 23.7 -53.4 -42.0 71.9 -36.9 24.2 60 53 L L T 45S- 0 0 99 2,-0.3 17,-0.3 -3,-0.3 -1,-0.3 0.424 104.1-123.1-107.8 -3.0 72.7 -39.3 21.4 61 54 L E T <5S+ 0 0 47 -4,-1.0 16,-1.4 1,-0.3 2,-0.2 0.720 82.7 80.1 68.9 20.9 73.5 -36.7 18.8 62 55 L A E < -GH 57 76B 2 -5,-1.1 -5,-1.2 14,-0.3 -2,-0.3 -0.829 64.3-136.7-146.1-176.1 70.9 -38.1 16.5 63 56 L A E -GH 56 75B 8 12,-2.2 12,-3.1 -2,-0.2 2,-0.3 -0.935 18.6-107.8-148.7 163.6 67.1 -38.1 15.8 64 57 L F E - H 0 74B 30 -9,-2.3 -10,-2.8 -2,-0.3 2,-0.4 -0.752 24.2-155.5-106.4 146.0 64.2 -40.4 14.9 65 58 L V E -GH 53 73B 0 8,-2.4 8,-2.6 -2,-0.3 2,-1.1 -0.943 16.4-147.9-124.3 131.9 62.3 -40.8 11.7 66 59 L D E +G 52 0B 54 -14,-2.7 -14,-1.7 -2,-0.4 6,-0.1 -0.843 29.6 162.5 -88.2 99.6 58.9 -42.0 10.8 67 60 L Y S S- 0 0 16 -2,-1.1 -17,-0.3 4,-0.2 -1,-0.2 0.567 71.5 -84.2 -94.9 -11.5 59.6 -43.4 7.4 68 61 L G S S+ 0 0 55 1,-0.4 4,-0.1 -3,-0.2 -2,-0.1 0.268 90.8 121.5 132.9 -5.4 56.4 -45.3 7.6 69 62 L A S S- 0 0 33 2,-0.3 -1,-0.4 1,-0.1 4,-0.1 -0.291 77.6-109.4 -63.1 161.3 57.1 -48.5 9.6 70 63 L E S S+ 0 0 114 2,-0.1 2,-0.4 1,-0.0 -1,-0.1 0.794 111.8 51.2 -58.5 -30.0 55.2 -49.2 12.8 71 64 L R S S- 0 0 91 42,-0.0 -2,-0.3 43,-0.0 -4,-0.2 -0.908 93.0-119.3-114.3 142.4 58.6 -48.4 14.4 72 65 L H - 0 0 65 -2,-0.4 -6,-0.2 -6,-0.1 43,-0.2 -0.451 24.5-121.1 -82.7 150.4 60.7 -45.3 13.7 73 66 L G E -H 65 0B 0 -8,-2.6 -8,-2.4 41,-0.2 2,-0.5 -0.375 20.5-116.7 -86.6 167.0 64.2 -45.3 12.3 74 67 L F E -Hi 64 116B 68 41,-3.6 43,-2.5 -10,-0.2 -10,-0.2 -0.901 23.7-169.7-111.8 127.4 67.3 -43.9 13.9 75 68 L L E -H 63 0B 12 -12,-3.1 -12,-2.2 -2,-0.5 43,-0.1 -0.835 11.5-149.9-126.8 97.9 69.2 -41.1 12.2 76 69 L P E > -H 62 0B 35 0, 0.0 3,-1.4 0, 0.0 4,-0.3 -0.128 22.3-118.7 -65.4 154.4 72.7 -40.1 13.6 77 70 L L G > S+ 0 0 49 -16,-1.4 3,-2.1 -17,-0.3 -15,-0.1 0.920 112.5 60.2 -58.4 -47.7 74.2 -36.6 13.5 78 71 L K G 3 S+ 0 0 86 1,-0.3 -1,-0.3 -17,-0.1 -16,-0.1 0.602 103.7 56.8 -57.3 -8.3 77.2 -37.6 11.3 79 72 L E G < S+ 0 0 50 -3,-1.4 40,-1.9 38,-0.1 41,-0.6 0.495 77.6 104.3-106.0 -6.8 74.5 -38.7 8.7 80 73 L I S < S- 0 0 38 -3,-2.1 2,-0.2 -4,-0.3 5,-0.1 -0.638 72.3-119.6 -81.2 132.9 72.5 -35.4 8.2 81 74 L A >> - 0 0 4 -2,-0.4 3,-3.1 1,-0.1 4,-0.7 -0.513 8.5-130.0 -77.7 134.3 73.3 -33.6 4.9 82 75 L R G >4 S+ 0 0 95 1,-0.3 3,-0.7 -2,-0.2 -1,-0.1 0.802 113.0 58.7 -47.8 -31.5 74.7 -30.0 4.9 83 76 L E G 34 S+ 0 0 98 1,-0.2 -1,-0.3 50,-0.2 -2,-0.0 0.675 101.7 52.7 -72.5 -20.5 71.8 -29.3 2.5 84 77 L Y G <4 S+ 0 0 30 -3,-3.1 -1,-0.2 1,-0.1 -2,-0.2 0.507 91.7 87.8 -92.2 -5.4 69.2 -30.5 5.0 85 78 L F << 0 0 38 -4,-0.7 10,-0.1 -3,-0.7 -1,-0.1 -0.804 360.0 360.0-101.0 98.9 70.6 -28.1 7.7 86 79 L P 0 0 149 0, 0.0 -3,-0.1 0, 0.0 -2,-0.1 0.037 360.0 360.0 -33.3 360.0 69.0 -24.6 7.5 87 ! 0 0 0 0, 0.0 0, 0.0 0, 0.0 0, 0.0 0.000 360.0 360.0 360.0 360.0 0.0 0.0 0.0 88 87 L R 0 0 154 0, 0.0 2,-0.1 0, 0.0 0, 0.0 0.000 360.0 360.0 360.0 132.1 80.5 -29.3 15.7 89 88 L P - 0 0 104 0, 0.0 2,-0.3 0, 0.0 3,-0.0 -0.358 360.0 -97.4-101.6 179.8 76.9 -28.5 16.7 90 89 L N > - 0 0 39 -2,-0.1 3,-1.5 1,-0.1 4,-0.4 -0.683 26.8-124.3 -93.0 155.5 74.1 -30.2 18.6 91 90 L I G >> S+ 0 0 18 1,-0.3 4,-1.3 -2,-0.3 3,-1.3 0.833 111.9 71.1 -67.7 -29.9 71.3 -32.0 16.7 92 91 L K G 34 S+ 0 0 63 1,-0.3 -1,-0.3 2,-0.1 -36,-0.0 0.666 99.7 51.6 -54.6 -13.8 68.9 -29.8 18.6 93 92 L D G <4 S+ 0 0 119 -3,-1.5 -1,-0.3 3,-0.0 -2,-0.2 0.681 110.4 41.5 -98.7 -23.0 70.4 -27.2 16.3 94 93 L V T <4 S+ 0 0 40 -3,-1.3 2,-0.3 -4,-0.4 -2,-0.1 0.914 108.4 36.6 -98.2 -52.7 70.0 -28.7 12.8 95 94 L L < - 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