==== Secondary Structure Definition by the program DSSP, updated CMBI version by ElmK / April 1,2000 ==== DATE=13-SEP-2013 . REFERENCE W. KABSCH AND C.SANDER, BIOPOLYMERS 22 (1983) 2577-2637 . HEADER RNA BINDING PROTEIN/IMMUNE SYSTEM 03-FEB-12 4DK3 . COMPND 2 MOLECULE: SINGLE DOMAIN ANTIBODY VHH; . SOURCE 2 ORGANISM_SCIENTIFIC: LAMA GLAMA; . AUTHOR Y.-J.PARK,W.HOL . 417 12 2 2 0 TOTAL NUMBER OF RESIDUES, NUMBER OF CHAINS, NUMBER OF SS-BRIDGES(TOTAL,INTRACHAIN,INTERCHAIN) . 21638.0 ACCESSIBLE SURFACE OF PROTEIN (ANGSTROM**2) . 293 70.3 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(J) , SAME NUMBER PER 100 RESIDUES . 19 4.6 TOTAL NUMBER OF HYDROGEN BONDS IN PARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 176 42.2 TOTAL NUMBER OF HYDROGEN BONDS IN ANTIPARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 5 1.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-5), SAME NUMBER PER 100 RESIDUES . 1 0.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-4), SAME NUMBER PER 100 RESIDUES . 3 0.7 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-3), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-2), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-1), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+0), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+1), SAME NUMBER PER 100 RESIDUES . 46 11.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+2), SAME NUMBER PER 100 RESIDUES . 26 6.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+3), SAME NUMBER PER 100 RESIDUES . 5 1.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+4), SAME NUMBER PER 100 RESIDUES . 2 0.5 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+5), SAME NUMBER PER 100 RESIDUES . 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 *** HISTOGRAMS OF *** . 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 RESIDUES PER ALPHA HELIX . 0 0 1 3 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PARALLEL BRIDGES PER LADDER . 1 4 8 2 2 10 2 2 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 ANTIPARALLEL BRIDGES PER LADDER . 0 0 3 0 0 2 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 LADDERS PER SHEET . # RESIDUE AA STRUCTURE BP1 BP2 ACC N-H-->O O-->H-N N-H-->O O-->H-N TCO KAPPA ALPHA PHI PSI X-CA Y-CA Z-CA 1 1 A Q 0 0 139 0, 0.0 26,-1.6 0, 0.0 2,-0.4 0.000 360.0 360.0 360.0 -90.4 -10.1 18.4 -25.3 2 2 A V E -A 26 0A 19 24,-0.2 2,-0.5 25,-0.1 24,-0.2 -0.996 360.0-163.4-132.4 129.0 -9.8 16.4 -28.5 3 3 A Q E -A 25 0A 115 22,-2.0 22,-1.8 -2,-0.4 2,-0.6 -0.949 2.5-162.8-110.9 132.4 -7.2 16.7 -31.3 4 4 A L E -A 24 0A 5 -2,-0.5 115,-0.7 113,-0.4 2,-0.5 -0.959 10.7-166.5-112.5 113.3 -7.8 15.2 -34.8 5 5 A Q E -A 23 0A 106 18,-3.0 18,-3.2 -2,-0.6 2,-0.3 -0.895 6.2-157.8-108.8 122.9 -4.5 14.9 -36.7 6 6 A E E +A 22 0A 21 -2,-0.5 2,-0.3 16,-0.2 16,-0.2 -0.711 18.8 154.3-106.4 146.8 -4.7 14.3 -40.4 7 7 A S E +A 21 0A 69 14,-1.9 14,-2.9 -2,-0.3 114,-0.1 -0.962 38.2 72.9-156.5 171.8 -2.2 12.8 -42.9 8 8 A G S S+ 0 0 35 -2,-0.3 11,-0.5 12,-0.2 12,-0.1 0.248 70.6 117.8 103.4 -9.8 -1.9 10.9 -46.2 9 9 A G + 0 0 28 12,-0.1 2,-0.3 9,-0.1 114,-0.2 0.179 25.4 104.9 -75.9-165.8 -2.8 13.8 -48.5 10 10 A G E -d 123 0B 30 112,-1.8 114,-2.6 -3,-0.1 2,-0.4 -0.900 66.4 -28.4 133.2-158.1 -0.8 15.6 -51.2 11 11 A L E +d 124 0B 124 -2,-0.3 2,-0.3 112,-0.2 114,-0.2 -0.849 56.0 156.7-104.7 131.9 -0.4 15.9 -55.0 12 12 A V E -d 125 0B 29 112,-2.1 114,-1.9 -2,-0.4 2,-0.1 -0.981 39.4-107.9-145.1 155.1 -1.3 13.0 -57.5 13 13 A Q E > -d 126 0B 149 -2,-0.3 3,-1.8 112,-0.2 73,-0.2 -0.454 54.6 -78.7 -77.5 157.4 -2.3 12.7 -61.1 14 14 A A T 3 S+ 0 0 47 112,-2.2 73,-0.2 1,-0.3 -1,-0.1 -0.254 121.4 29.6 -58.9 142.4 -5.9 11.7 -62.1 15 15 A G T 3 S+ 0 0 58 71,-3.0 70,-0.5 1,-0.3 -1,-0.3 0.317 100.1 122.8 83.7 -6.9 -6.7 8.0 -61.7 16 16 A G < - 0 0 22 -3,-1.8 70,-2.0 70,-0.3 2,-0.3 -0.216 53.5-131.7 -79.6 173.8 -4.1 8.0 -58.9 17 17 A S + 0 0 71 68,-0.1 2,-0.3 67,-0.1 67,-0.2 -0.962 20.4 178.9-135.6 148.7 -4.7 7.0 -55.3 18 18 A L E - B 0 83A 24 65,-1.8 65,-2.5 -2,-0.3 2,-0.4 -0.922 20.5-140.8-135.7 163.9 -4.1 8.2 -51.8 19 19 A R E - B 0 82A 134 -11,-0.5 63,-0.2 -2,-0.3 2,-0.2 -0.892 19.6-158.3-129.8 100.8 -4.9 6.9 -48.3 20 20 A L E - B 0 81A 0 61,-2.2 61,-2.3 -2,-0.4 2,-0.4 -0.513 7.0-162.9 -77.8 147.4 -6.0 9.6 -45.8 21 21 A S E -AB 7 80A 31 -14,-2.9 -14,-1.9 59,-0.2 2,-0.6 -0.993 11.1-164.0-138.1 125.7 -5.7 9.0 -42.1 22 22 A a E -AB 6 79A 0 57,-2.8 57,-1.7 -2,-0.4 2,-0.5 -0.947 12.1-165.3-108.8 114.4 -7.3 10.8 -39.1 23 23 A A E -AB 5 78A 38 -18,-3.2 -18,-3.0 -2,-0.6 2,-0.4 -0.873 6.2-153.4-102.5 128.5 -5.7 10.1 -35.8 24 24 A A E +A 4 0A 14 53,-2.4 2,-0.3 -2,-0.5 -20,-0.2 -0.807 18.5 166.9-103.7 141.0 -7.5 11.0 -32.6 25 25 A S E +A 3 0A 53 -22,-1.8 -22,-2.0 -2,-0.4 2,-0.4 -0.917 44.0 46.1-142.6 167.9 -6.0 11.8 -29.2 26 26 A G E S-A 2 0A 62 -2,-0.3 2,-1.5 -24,-0.2 -24,-0.2 -0.841 119.1 -2.0 96.9-138.1 -7.3 13.4 -26.0 27 27 A R > + 0 0 102 -26,-1.6 3,-1.5 -2,-0.4 4,-0.2 -0.544 69.9 161.5 -90.5 72.3 -10.6 12.1 -24.7 28 28 A T G > + 0 0 27 -2,-1.5 3,-1.2 1,-0.3 -1,-0.2 0.633 61.3 80.6 -67.3 -12.2 -11.4 9.7 -27.5 29 29 A S G 3 S+ 0 0 37 1,-0.3 381,-0.3 -3,-0.2 -1,-0.3 0.781 90.0 51.6 -64.3 -28.1 -13.9 8.0 -25.1 30 30 A S G < S+ 0 0 17 -3,-1.5 2,-0.3 379,-0.1 -1,-0.3 0.487 90.0 99.9 -88.0 -5.4 -16.5 10.7 -25.9 31 31 A L < + 0 0 5 -3,-1.2 69,-0.2 -4,-0.2 3,-0.1 -0.663 35.2 151.1 -88.4 138.0 -16.1 10.3 -29.7 32 32 A Y + 0 0 30 67,-2.9 2,-0.2 1,-0.5 68,-0.1 0.512 56.0 39.6-126.2 -87.7 -18.7 8.2 -31.7 33 33 A S - 0 0 0 68,-0.3 66,-2.7 66,-0.2 -1,-0.5 -0.527 66.4-164.0 -70.8 137.0 -19.5 9.0 -35.3 34 34 A M E +EF 51 98B 8 17,-2.2 17,-3.1 64,-0.2 2,-0.3 -0.932 10.9 179.4-120.4 147.8 -16.5 10.0 -37.4 35 35 A G E -EF 50 97B 0 62,-2.4 62,-2.6 -2,-0.3 2,-0.5 -0.983 26.3-133.1-149.4 157.2 -16.6 11.7 -40.8 36 36 A W E +EF 49 96B 0 13,-4.5 12,-2.5 -2,-0.3 13,-1.5 -0.953 27.3 176.8-112.5 130.5 -14.6 13.1 -43.7 37 37 A F E -EF 47 95B 0 58,-3.6 58,-3.5 -2,-0.5 2,-0.3 -0.875 9.4-157.5-126.7 158.7 -15.5 16.5 -45.2 38 38 A R E -EF 46 94B 9 8,-2.5 8,-3.0 -2,-0.3 2,-0.4 -0.996 6.5-163.3-142.3 150.2 -13.9 18.6 -47.9 39 39 A Q E - F 0 93B 55 54,-2.3 54,-2.4 -2,-0.3 6,-0.1 -0.953 3.8-162.8-136.6 115.2 -13.7 22.2 -49.0 40 40 A A - 0 0 19 -2,-0.4 3,-0.4 4,-0.2 52,-0.1 -0.678 47.5 -82.5 -86.4 149.4 -12.7 23.5 -52.4 41 41 A P S S+ 0 0 121 0, 0.0 2,-0.9 0, 0.0 -1,-0.1 -0.240 114.4 11.6 -57.7 137.8 -11.7 27.3 -52.5 42 42 A G S S+ 0 0 92 1,-0.3 -3,-0.0 -3,-0.1 0, 0.0 -0.399 117.2 80.1 91.6 -53.8 -14.7 29.6 -52.9 43 43 A K S S- 0 0 151 -2,-0.9 -1,-0.3 -3,-0.4 2,-0.3 -0.064 80.9-110.8 -76.3 178.0 -17.2 26.8 -52.1 44 44 A E - 0 0 157 -3,-0.1 -4,-0.2 1,-0.1 -1,-0.1 -0.791 44.1 -64.7-116.2 162.1 -18.1 25.4 -48.7 45 45 A R - 0 0 78 -2,-0.3 2,-0.4 -6,-0.1 -6,-0.2 -0.096 50.2-165.1 -43.6 118.9 -17.6 22.2 -46.8 46 46 A E E -E 38 0B 76 -8,-3.0 -8,-2.5 -3,-0.0 2,-0.4 -0.867 25.9-108.2-107.3 147.0 -19.3 19.2 -48.4 47 47 A F E +E 37 0B 26 -2,-0.4 -10,-0.2 -10,-0.2 3,-0.1 -0.576 38.3 171.2 -68.4 123.8 -19.9 15.8 -46.7 48 48 A V E - 0 0 0 -12,-2.5 13,-2.4 -2,-0.4 2,-0.3 0.847 58.5 -25.2 -95.3 -67.9 -17.6 13.1 -48.2 49 49 A A E -EG 36 60B 0 -13,-1.5 -13,-4.5 11,-0.2 -1,-0.3 -0.990 53.1-171.1-150.6 150.2 -18.0 10.0 -46.0 50 50 A A E -EG 35 59B 0 9,-1.7 9,-1.9 -2,-0.3 2,-0.3 -0.985 5.1-158.9-145.8 156.0 -18.9 9.1 -42.5 51 51 A I E -EG 34 58B 4 -17,-3.1 -17,-2.2 -2,-0.3 57,-0.2 -0.991 20.5-116.8-140.4 140.0 -18.9 5.9 -40.3 52 52 A S > - 0 0 0 5,-2.4 3,-0.8 -2,-0.3 53,-0.1 -0.188 36.5-104.6 -67.0 166.3 -20.6 4.8 -37.1 53 53 A R T 3 S+ 0 0 30 50,-0.4 52,-0.2 53,-0.3 51,-0.1 0.891 122.5 47.4 -61.4 -41.9 -18.8 4.0 -33.9 54 54 A N T 3 S- 0 0 63 50,-2.3 -1,-0.2 1,-0.1 51,-0.1 0.551 108.5-125.5 -76.7 -8.3 -19.1 0.2 -34.4 55 55 A G S < S+ 0 0 30 -3,-0.8 -2,-0.1 49,-0.3 50,-0.1 0.335 82.4 111.9 81.3 -7.2 -17.9 0.5 -38.0 56 56 A A + 0 0 67 1,-0.1 2,-0.5 -5,-0.1 49,-0.1 0.718 68.4 56.3 -70.8 -23.0 -21.0 -1.3 -39.2 57 57 A N + 0 0 30 49,-0.0 -5,-2.4 2,-0.0 2,-0.3 -0.948 59.5 173.5-126.1 127.6 -22.6 1.7 -40.9 58 58 A T E -G 51 0B 61 -2,-0.5 2,-0.4 -7,-0.2 -7,-0.2 -0.931 10.4-165.7-130.2 151.0 -21.2 4.0 -43.6 59 59 A Y E -G 50 0B 72 -9,-1.9 -9,-1.7 -2,-0.3 2,-0.4 -0.996 8.0-169.4-139.9 132.4 -22.6 6.8 -45.8 60 60 A Y E -G 49 0B 41 -2,-0.4 -11,-0.2 -11,-0.2 2,-0.1 -0.928 27.8-115.1-123.6 143.4 -21.1 8.4 -48.9 61 61 A T >> - 0 0 37 -13,-2.4 3,-1.7 -2,-0.4 4,-0.6 -0.506 42.2-111.4 -66.4 145.1 -21.9 11.4 -51.0 62 62 A D G >4 S+ 0 0 122 1,-0.3 3,-1.1 2,-0.2 -1,-0.1 0.839 111.5 58.7 -52.1 -42.3 -23.0 10.2 -54.5 63 63 A S G 34 S+ 0 0 58 1,-0.2 -1,-0.3 3,-0.1 -15,-0.0 0.651 108.7 44.1 -66.8 -17.3 -19.9 11.4 -56.4 64 64 A V G X> S+ 0 0 0 -3,-1.7 4,-2.6 -16,-0.2 3,-1.2 0.453 78.9 127.0-108.0 -1.7 -17.5 9.3 -54.4 65 65 A K T << S+ 0 0 119 -3,-1.1 4,-0.1 -4,-0.6 -3,-0.0 -0.362 82.7 9.4 -59.8 128.0 -19.4 6.1 -54.2 66 66 A G T 34 S+ 0 0 81 2,-0.5 -1,-0.2 -2,-0.1 3,-0.1 -0.010 125.6 64.4 91.4 -28.1 -17.2 3.2 -55.4 67 67 A R T <4 S+ 0 0 28 -3,-1.2 17,-2.5 1,-0.1 2,-0.4 0.860 99.4 45.6 -94.9 -43.4 -14.1 5.3 -55.5 68 68 A F E < -C 83 0A 3 -4,-2.6 -2,-0.5 15,-0.2 2,-0.4 -0.837 64.9-165.6-104.7 143.1 -13.6 6.2 -51.8 69 69 A T E -C 82 0A 70 13,-2.4 13,-3.2 -2,-0.4 2,-0.5 -0.992 5.5-153.6-129.2 132.1 -13.9 3.7 -48.9 70 70 A I E +C 81 0A 7 -2,-0.4 2,-0.3 11,-0.2 11,-0.2 -0.893 23.1 155.9-107.9 131.5 -14.2 4.6 -45.2 71 71 A S E -C 80 0A 60 9,-1.4 9,-3.1 -2,-0.5 2,-0.3 -0.931 20.0-152.4-141.2 167.8 -13.0 2.4 -42.3 72 72 A R E -C 79 0A 43 -2,-0.3 2,-0.5 7,-0.3 7,-0.2 -0.987 12.6-155.2-144.2 152.5 -11.9 2.8 -38.7 73 73 A D E > -C 78 0A 69 5,-2.7 5,-2.4 -2,-0.3 4,-0.4 -0.927 10.5-174.0-127.0 102.1 -9.7 1.1 -36.1 74 74 A N T > 5S+ 0 0 38 -2,-0.5 3,-0.7 1,-0.2 -1,-0.1 0.870 83.7 58.8 -68.4 -36.8 -11.0 1.9 -32.7 75 75 A A T 3 5S+ 0 0 86 1,-0.3 -1,-0.2 2,-0.1 -2,-0.0 0.795 116.4 35.8 -60.5 -31.2 -8.1 0.2 -30.9 76 76 A K T 3 5S- 0 0 149 -3,-0.4 -1,-0.3 2,-0.2 -2,-0.2 0.428 108.6-130.5 -95.8 -3.3 -5.7 2.6 -32.8 77 77 A N T < 5 + 0 0 51 -3,-0.7 -53,-2.4 -4,-0.4 2,-0.3 0.867 63.8 128.4 50.9 43.6 -8.3 5.4 -32.5 78 78 A T E < -BC 23 73A 28 -5,-2.4 -5,-2.7 -55,-0.2 2,-0.4 -0.887 45.4-160.0-125.4 154.4 -8.0 6.1 -36.3 79 79 A V E -BC 22 72A 0 -57,-1.7 -57,-2.8 -2,-0.3 2,-0.3 -0.994 16.4-152.2-129.7 141.4 -10.3 6.4 -39.3 80 80 A E E -BC 21 71A 60 -9,-3.1 -9,-1.4 -2,-0.4 2,-0.5 -0.847 13.4-155.4-114.8 149.1 -9.2 6.0 -42.9 81 81 A L E -BC 20 70A 0 -61,-2.3 -61,-2.2 -2,-0.3 2,-0.7 -0.892 13.7-158.2-120.5 99.2 -10.3 7.4 -46.3 82 82 A Q E -BC 19 69A 49 -13,-3.2 -13,-2.4 -2,-0.5 2,-0.7 -0.672 13.2-165.3 -72.1 109.1 -9.3 5.0 -49.1 83 83 A M E +BC 18 68A 0 -65,-2.5 -65,-1.8 -2,-0.7 2,-0.2 -0.872 14.5 167.5-108.7 104.7 -9.2 7.2 -52.2 84 84 A N + 0 0 69 -17,-2.5 -68,-0.1 -2,-0.7 -67,-0.1 -0.715 59.0 19.3-105.7 162.5 -9.1 5.4 -55.5 85 85 A S S S- 0 0 72 -70,-0.5 -1,-0.2 -2,-0.2 -68,-0.1 0.875 81.0-159.1 47.0 52.5 -9.7 6.7 -59.0 86 86 A L - 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