==== Secondary Structure Definition by the program DSSP, updated CMBI version by ElmK / April 1,2000 ==== DATE=20-JUL-2011 . REFERENCE W. KABSCH AND C.SANDER, BIOPOLYMERS 22 (1983) 2577-2637 . HEADER LIGASE 12-JUL-06 2DTH . COMPND 2 MOLECULE: 235AA LONG HYPOTHETICAL BIOTIN--[ACETYL-COA-CARBO . SOURCE 2 ORGANISM_SCIENTIFIC: PYROCOCCUS HORIKOSHII; . AUTHOR B.BAGAUTDINOV,N.KUNISHIMA,RIKEN STRUCTURAL GENOMICS/PROTEOMI . 470 2 0 0 0 TOTAL NUMBER OF RESIDUES, NUMBER OF CHAINS, NUMBER OF SS-BRIDGES(TOTAL,INTRACHAIN,INTERCHAIN) . 20841.0 ACCESSIBLE SURFACE OF PROTEIN (ANGSTROM**2) . 333 70.9 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(J) , SAME NUMBER PER 100 RESIDUES . 20 4.3 TOTAL NUMBER OF HYDROGEN BONDS IN PARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 117 24.9 TOTAL NUMBER OF HYDROGEN BONDS IN ANTIPARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-5), SAME NUMBER PER 100 RESIDUES . 8 1.7 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-4), SAME NUMBER PER 100 RESIDUES . 4 0.9 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-3), SAME NUMBER PER 100 RESIDUES . 2 0.4 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-2), SAME NUMBER PER 100 RESIDUES . 2 0.4 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-1), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+0), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+1), SAME NUMBER PER 100 RESIDUES . 38 8.1 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+2), SAME NUMBER PER 100 RESIDUES . 43 9.1 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+3), SAME NUMBER PER 100 RESIDUES . 94 20.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+4), SAME NUMBER PER 100 RESIDUES . 4 0.9 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+5), SAME NUMBER PER 100 RESIDUES . 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 *** HISTOGRAMS OF *** . 0 0 0 0 2 2 1 0 1 0 0 0 0 0 2 0 0 0 0 0 2 0 0 0 0 0 0 0 0 0 RESIDUES PER ALPHA HELIX . 2 0 0 0 0 2 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PARALLEL BRIDGES PER LADDER . 3 0 4 8 3 0 3 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 ANTIPARALLEL BRIDGES PER LADDER . 0 0 0 2 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 LADDERS PER SHEET . # RESIDUE AA STRUCTURE BP1 BP2 ACC N-H-->O O-->H-N N-H-->O O-->H-N TCO KAPPA ALPHA PHI PSI X-CA Y-CA Z-CA 1 1 A M 0 0 6 0, 0.0 389,-0.2 0, 0.0 388,-0.1 0.000 360.0 360.0 360.0 -7.6 40.9 27.2 36.8 2 2 A L - 0 0 5 250,-0.1 251,-0.1 387,-0.1 388,-0.1 0.717 360.0-142.6 -77.5 -20.7 39.9 26.7 33.1 3 3 A G + 0 0 21 1,-0.2 2,-0.2 250,-0.1 250,-0.1 0.812 33.3 168.5 65.1 34.7 43.5 25.9 32.3 4 4 A L - 0 0 4 6,-0.2 -1,-0.2 1,-0.1 272,-0.1 -0.537 20.5-171.4 -77.9 146.6 43.7 27.6 28.9 5 5 A K + 0 0 168 -2,-0.2 -1,-0.1 0, 0.0 6,-0.1 0.145 36.4 137.3-123.5 17.7 47.2 28.0 27.4 6 6 A T - 0 0 9 4,-0.2 156,-0.1 3,-0.2 247,-0.1 -0.143 51.1-140.3 -64.9 157.7 46.3 30.2 24.5 7 7 A S S S- 0 0 74 2,-0.1 -1,-0.1 4,-0.0 3,-0.1 0.816 87.8 -6.3 -90.8 -34.4 48.4 33.2 23.4 8 8 A I S > S+ 0 0 21 1,-0.3 3,-1.2 26,-0.1 2,-0.5 0.633 119.0 62.1-124.0 -66.2 45.6 35.6 22.5 9 9 A I T 3 S+ 0 0 2 1,-0.2 -1,-0.3 152,-0.1 26,-0.2 -0.546 112.9 17.2 -72.9 118.6 42.1 34.2 22.5 10 10 A G T 3 S+ 0 0 0 24,-3.0 -1,-0.2 -2,-0.5 -4,-0.2 0.666 75.0 124.8 98.0 15.9 41.1 33.1 26.0 11 11 A R S < S+ 0 0 74 -3,-1.2 242,-2.2 1,-0.3 2,-0.3 0.829 91.3 22.8 -72.3 -29.2 43.8 34.9 28.1 12 12 A R E S-A 252 0A 97 22,-0.3 24,-2.2 -4,-0.3 2,-0.4 -0.988 73.3-171.5-134.6 142.8 40.6 36.2 29.8 13 13 A V E -Ab 251 36A 3 238,-2.1 238,-3.2 -2,-0.3 2,-0.5 -0.999 7.4-159.4-137.8 131.0 37.1 34.7 29.9 14 14 A I E -Ab 250 37A 23 22,-3.5 24,-2.7 -2,-0.4 2,-0.4 -0.956 14.5-164.1-113.3 123.2 34.0 36.4 31.3 15 15 A Y E -Ab 249 38A 3 234,-3.1 234,-2.1 -2,-0.5 2,-0.4 -0.904 8.5-172.7-114.2 136.8 31.2 34.0 32.2 16 16 A F E -Ab 248 39A 13 22,-2.2 24,-1.6 -2,-0.4 25,-0.3 -0.978 24.7-145.7-124.5 136.6 27.5 34.8 32.9 17 17 A Q E S+ 0 0 39 230,-2.1 24,-0.6 -2,-0.4 2,-0.3 0.901 93.9 4.4 -63.5 -38.6 24.8 32.4 34.2 18 18 A E E S+ b 0 41A 108 229,-0.4 2,-0.3 22,-0.2 -1,-0.3 -0.998 72.4 164.8-148.4 145.1 22.4 34.4 32.0 19 19 A I E - b 0 42A 16 22,-1.5 24,-2.3 -2,-0.3 25,-0.4 -0.983 42.6-122.2-155.5 160.3 22.7 37.2 29.4 20 20 A T S S- 0 0 78 1,-0.3 24,-2.7 -2,-0.3 2,-0.3 0.981 95.4 -24.5 -69.0 -55.6 20.8 38.9 26.7 21 21 A S > - 0 0 4 22,-0.1 4,-1.5 21,-0.1 -1,-0.3 -0.939 45.5-145.3-159.6 133.3 23.5 38.2 24.1 22 22 A T H > S+ 0 0 4 -2,-0.3 4,-2.0 1,-0.2 41,-0.1 0.822 104.8 57.9 -70.0 -29.2 27.2 37.6 24.4 23 23 A N H > S+ 0 0 7 1,-0.2 4,-0.6 2,-0.2 -1,-0.2 0.864 106.2 48.6 -68.3 -34.5 27.7 39.4 21.1 24 24 A E H > S+ 0 0 74 2,-0.2 4,-1.5 1,-0.2 3,-0.4 0.838 112.0 47.6 -74.2 -32.9 26.0 42.5 22.5 25 25 A F H < S+ 0 0 58 -4,-1.5 4,-0.3 1,-0.2 -2,-0.2 0.872 104.9 60.7 -75.3 -34.3 28.1 42.5 25.7 26 26 A A H < S+ 0 0 4 -4,-2.0 -1,-0.2 -5,-0.2 -2,-0.2 0.694 113.3 37.5 -64.6 -19.0 31.3 42.0 23.7 27 27 A K H < S+ 0 0 48 -4,-0.6 -2,-0.2 -3,-0.4 -1,-0.1 0.910 110.4 50.7 -92.9 -69.1 30.7 45.3 21.9 28 28 A T S < S+ 0 0 100 -4,-1.5 2,-0.4 2,-0.1 -2,-0.1 0.363 105.3 83.7 -55.7 7.2 29.2 47.8 24.3 29 29 A S S S- 0 0 19 -4,-0.3 2,-0.5 -5,-0.1 -4,-0.0 -0.940 80.5-128.3-124.2 140.2 32.1 46.9 26.5 30 30 A Y + 0 0 210 -2,-0.4 2,-0.3 2,-0.0 -3,-0.1 -0.707 49.4 157.2 -76.4 126.0 35.8 48.0 26.7 31 31 A L - 0 0 30 -2,-0.5 2,-0.2 -4,-0.1 3,-0.1 -0.986 43.8-105.9-156.1 147.0 37.6 44.7 26.5 32 32 A E > - 0 0 80 -2,-0.3 3,-2.1 1,-0.1 33,-0.2 -0.480 50.4 -90.9 -74.8 144.6 41.1 43.5 25.6 33 33 A E T 3 S+ 0 0 43 1,-0.3 33,-0.2 -2,-0.2 -1,-0.1 -0.209 114.2 31.2 -52.0 142.5 41.5 41.6 22.3 34 34 A G T 3 S+ 0 0 0 31,-3.4 -24,-3.0 1,-0.3 -22,-0.3 0.232 82.5 141.3 90.8 -14.9 41.1 37.8 22.8 35 35 A T < - 0 0 3 -3,-2.1 30,-2.4 -26,-0.2 2,-0.5 -0.418 40.8-146.9 -65.0 134.6 38.7 38.2 25.7 36 36 A V E -bC 13 64A 0 -24,-2.2 -22,-3.5 28,-0.2 2,-0.5 -0.906 8.9-160.4-110.1 130.3 35.9 35.6 25.6 37 37 A I E +bC 14 63A 0 26,-3.1 26,-2.4 -2,-0.5 2,-0.3 -0.925 20.0 171.1-107.2 126.6 32.4 36.1 26.7 38 38 A V E -bC 15 62A 0 -24,-2.7 -22,-2.2 -2,-0.5 2,-0.3 -0.986 9.9-175.2-138.4 146.8 30.4 33.0 27.5 39 39 A A E -bC 16 61A 0 22,-1.7 22,-0.6 -2,-0.3 -22,-0.2 -0.997 32.9-132.2-143.0 147.7 27.0 32.3 29.0 40 40 A D E S+ 0 0 12 -24,-1.6 19,-0.5 -2,-0.3 2,-0.3 0.871 96.4 3.8 -63.1 -38.2 24.9 29.2 30.0 41 41 A K E -b 18 0A 87 -24,-0.6 -22,-1.5 -25,-0.3 2,-0.4 -0.992 62.5-147.8-148.9 152.6 21.9 30.8 28.3 42 42 A Q E -b 19 0A 22 13,-1.6 13,-0.5 -2,-0.3 3,-0.3 -0.971 1.7-165.0-120.5 137.4 21.0 33.8 26.2 43 43 A T S S+ 0 0 63 -24,-2.3 -23,-0.2 -2,-0.4 -22,-0.1 0.650 94.1 28.4 -93.4 -18.5 17.6 35.4 26.3 44 44 A M S S+ 0 0 119 -24,-2.7 -1,-0.2 -25,-0.4 2,-0.1 -0.242 79.8 157.1-136.0 45.0 18.1 37.4 23.2 45 45 A G - 0 0 2 8,-0.3 8,-2.6 -3,-0.3 2,-0.3 -0.460 18.3-178.2 -71.6 144.3 20.5 35.4 21.1 46 46 A H B -Q 52 0B 46 6,-0.3 6,-0.2 -2,-0.1 2,-0.1 -0.967 13.7-138.5-143.7 162.5 20.4 36.1 17.4 47 47 A G > - 0 0 16 4,-3.2 3,-1.7 -2,-0.3 6,-0.1 -0.300 42.4 -67.5-108.5-165.4 21.9 35.0 14.1 48 48 A R T 3 S+ 0 0 39 1,-0.3 68,-0.0 2,-0.2 -2,-0.0 0.823 129.3 37.3 -48.3 -54.9 23.3 36.3 10.8 49 49 A L T 3 S- 0 0 52 3,-0.1 -1,-0.3 180,-0.1 3,-0.1 0.160 123.6 -99.7 -92.0 19.2 20.1 37.6 9.3 50 50 A N S < S+ 0 0 130 -3,-1.7 -2,-0.2 1,-0.2 2,-0.1 0.654 77.9 142.9 71.2 22.3 18.9 38.8 12.7 51 51 A R - 0 0 191 1,-0.1 -4,-3.2 -5,-0.0 -1,-0.2 -0.474 61.3 -85.2 -83.9 160.8 16.7 35.8 13.2 52 52 A K B -Q 46 0B 115 -6,-0.2 2,-0.6 -2,-0.1 -6,-0.3 -0.397 34.2-149.8 -71.2 143.5 16.5 34.4 16.8 53 53 A W - 0 0 39 -8,-2.6 -8,-0.3 -2,-0.1 2,-0.3 -0.948 18.6-130.2-115.5 111.0 19.2 31.9 17.9 54 54 A E + 0 0 92 -2,-0.6 -11,-0.1 1,-0.1 -8,-0.1 -0.436 29.7 178.9 -61.2 121.2 17.8 29.5 20.5 55 55 A S + 0 0 7 -13,-0.5 -13,-1.6 -2,-0.3 -1,-0.1 -0.483 11.5 159.3-126.7 62.8 20.3 29.5 23.4 56 56 A P > - 0 0 50 0, 0.0 3,-1.2 0, 0.0 78,-0.1 -0.282 53.7 -71.9 -80.2 169.6 19.1 27.1 26.1 57 57 A E T 3 S+ 0 0 114 1,-0.2 78,-0.3 76,-0.1 -16,-0.1 -0.287 114.7 48.4 -56.3 143.2 21.3 25.6 28.9 58 58 A G T 3 S+ 0 0 10 1,-0.4 75,-3.5 -18,-0.2 76,-0.6 0.225 91.3 89.1 110.7 -15.7 23.6 22.9 27.5 59 59 A G S < S- 0 0 0 -3,-1.2 2,-0.7 -19,-0.5 -1,-0.4 -0.563 78.3-108.5-109.0 176.2 25.0 24.6 24.5 60 60 A L E + D 0 130A 2 70,-2.8 70,-2.7 -2,-0.2 2,-0.6 -0.921 36.0 174.1-110.0 111.3 28.1 26.8 24.0 61 61 A W E +CD 39 129A 1 -2,-0.7 -22,-1.7 -22,-0.6 2,-0.3 -0.954 19.0 148.7-118.3 109.0 27.3 30.4 23.4 62 62 A L E -CD 38 128A 2 66,-2.4 66,-2.8 -2,-0.6 2,-0.3 -0.942 31.1-156.7-142.1 163.6 30.3 32.7 23.2 63 63 A S E -CD 37 127A 0 -26,-2.4 -26,-3.1 -2,-0.3 2,-0.5 -0.996 10.6-151.0-140.4 141.6 31.8 35.8 21.7 64 64 A I E -CD 36 126A 0 62,-2.1 62,-2.7 -2,-0.3 2,-0.4 -0.962 12.8-136.3-120.3 128.4 35.4 36.8 21.2 65 65 A V E + D 0 125A 3 -30,-2.4 -31,-3.4 -2,-0.5 2,-0.3 -0.688 34.5 172.2 -81.1 128.4 36.6 40.4 21.1 66 66 A L E + D 0 124A 0 58,-3.0 58,-3.7 -2,-0.4 -33,-0.0 -0.978 31.3 180.0-140.1 152.6 39.1 41.0 18.3 67 67 A S - 0 0 23 -2,-0.3 2,-0.1 56,-0.2 56,-0.1 -0.493 24.0-166.8-147.5 65.8 40.9 43.8 16.6 68 68 A P - 0 0 14 0, 0.0 2,-1.4 0, 0.0 -2,-0.1 -0.375 16.4-142.9 -61.4 126.6 42.9 42.1 13.8 69 69 A K + 0 0 181 -2,-0.1 53,-0.0 3,-0.0 -2,-0.0 -0.745 58.1 118.9 -92.9 87.3 45.6 44.4 12.4 70 70 A V S S- 0 0 19 -2,-1.4 5,-0.1 104,-0.0 102,-0.1 -0.962 75.1 -72.0-144.4 157.0 45.3 43.2 8.8 71 71 A P > - 0 0 80 0, 0.0 3,-2.7 0, 0.0 4,-0.5 -0.231 44.4-118.7 -54.9 139.8 44.3 45.0 5.6 72 72 A Q G > S+ 0 0 138 1,-0.3 3,-0.5 2,-0.2 -3,-0.0 0.699 110.6 68.8 -51.6 -23.5 40.6 45.9 5.4 73 73 A K G 3 S+ 0 0 152 1,-0.2 -1,-0.3 2,-0.1 4,-0.0 0.643 101.7 45.0 -73.0 -15.0 40.3 43.7 2.3 74 74 A D G X S+ 0 0 23 -3,-2.7 3,-1.4 1,-0.1 -1,-0.2 0.462 84.9 91.9-107.2 -4.6 40.9 40.5 4.4 75 75 A L G X S+ 0 0 28 -3,-0.5 3,-2.5 -4,-0.5 4,-0.3 0.872 75.7 65.7 -58.6 -40.5 38.6 41.3 7.3 76 76 A P G > S+ 0 0 69 0, 0.0 3,-0.8 0, 0.0 4,-0.3 0.717 88.7 69.3 -56.7 -19.8 35.6 39.5 5.7 77 77 A K G X> S+ 0 0 0 -3,-1.4 4,-0.9 1,-0.2 3,-0.5 0.566 76.6 85.9 -75.7 -6.9 37.5 36.2 6.1 78 78 A I H X> S+ 0 0 1 -3,-2.5 4,-1.0 1,-0.2 3,-0.6 0.861 80.1 59.6 -60.7 -38.8 37.1 36.5 9.9 79 79 A V H <> S+ 0 0 4 -3,-0.8 4,-1.4 -4,-0.3 -1,-0.2 0.831 100.4 56.2 -62.5 -31.1 33.7 34.8 9.7 80 80 A F H <> S+ 0 0 3 -3,-0.5 4,-3.1 -4,-0.3 5,-0.3 0.841 95.2 68.0 -68.9 -31.8 35.3 31.7 8.2 81 81 A L H <>S+ 0 0 0 -4,-2.8 5,-2.6 1,-0.2 3,-1.6 0.919 105.4 57.1 -58.9 -42.7 34.0 18.6 16.0 91 91 A K H ><5S+ 0 0 105 -4,-2.3 3,-1.8 1,-0.3 -1,-0.2 0.895 102.1 55.9 -52.9 -43.5 36.7 16.4 14.3 92 92 A E H 3<5S+ 0 0 112 -4,-1.5 -1,-0.3 1,-0.3 -2,-0.2 0.709 107.7 48.6 -63.4 -20.5 38.3 16.1 17.7 93 93 A F T <<5S- 0 0 41 -3,-1.6 -1,-0.3 -4,-0.8 -2,-0.2 -0.007 124.5-103.9-108.9 28.2 35.0 14.8 19.0 94 94 A S T < 5S+ 0 0 106 -3,-1.8 2,-0.4 1,-0.2 -3,-0.2 0.730 80.4 131.3 56.2 31.7 34.7 12.3 16.2 95 95 A I < - 0 0 8 -5,-2.6 2,-1.0 -6,-0.1 -2,-0.2 -0.927 51.6-143.4-115.1 135.1 32.1 14.4 14.4 96 96 A D + 0 0 115 -2,-0.4 12,-0.4 -3,-0.1 2,-0.3 -0.764 28.1 174.9-100.2 91.5 32.3 15.2 10.7 97 97 A G - 0 0 0 -2,-1.0 2,-0.3 -10,-0.3 10,-0.2 -0.691 11.6-160.9 -96.6 151.1 31.0 18.7 10.2 98 98 A R E -E 106 0A 94 8,-2.8 8,-3.1 -2,-0.3 2,-0.4 -0.930 21.6-109.8-128.4 153.6 31.0 20.7 7.0 99 99 A I E -Ef 105 186A 0 86,-3.4 88,-2.9 -2,-0.3 2,-0.7 -0.691 15.7-157.1 -90.8 131.5 30.6 24.4 6.3 100 100 A K E > -E 104 0A 43 4,-2.8 4,-2.0 -2,-0.4 3,-0.3 -0.928 54.7 -71.7-103.0 110.1 27.5 25.9 4.6 101 101 A W T 4 S+ 0 0 7 -2,-0.7 131,-0.1 86,-0.3 2,-0.1 -0.142 105.9 1.0 -45.2 135.1 28.7 29.2 3.1 102 102 A P T 4 S- 0 0 1 0, 0.0 -1,-0.3 0, 0.0 -19,-0.2 -0.956 133.0 -14.4-105.8 -18.3 29.4 31.4 4.7 103 103 A N T 4 S+ 0 0 3 -3,-0.3 12,-0.4 -2,-0.1 2,-0.2 0.455 93.1 102.2-123.4 -5.7 29.1 30.8 8.4 104 104 A D E < -E 100 0A 4 -4,-2.0 -4,-2.8 10,-0.2 2,-0.5 -0.536 48.3-154.7 -89.5 153.1 27.1 27.7 9.1 105 105 A V E -EG 99 113A 0 8,-1.9 7,-2.4 -6,-0.3 8,-1.4 -0.993 18.0-174.3-125.7 123.9 28.3 24.3 10.2 106 106 A L E -EG 98 111A 11 -8,-3.1 -8,-2.8 -2,-0.5 2,-0.5 -0.833 21.0-155.3-119.2 155.1 26.2 21.2 9.4 107 107 A V E > S- G 0 110A 9 3,-2.3 3,-1.4 -2,-0.3 -10,-0.1 -0.973 92.5 -15.7-128.9 110.6 26.4 17.5 10.2 108 108 A N T 3 S- 0 0 131 -2,-0.5 -1,-0.1 -12,-0.4 3,-0.1 0.928 130.7 -55.2 56.7 44.3 24.5 15.5 7.5 109 109 A Y T 3 S+ 0 0 104 1,-0.2 2,-0.5 -11,-0.0 -1,-0.3 0.556 113.1 120.6 66.1 11.8 22.9 18.8 6.5 110 110 A K E < -G 107 0A 81 -3,-1.4 -3,-2.3 30,-0.0 2,-0.4 -0.915 69.8-113.8-108.1 130.1 21.6 19.5 10.0 111 111 A K E +G 106 0A 67 -2,-0.5 30,-2.3 28,-0.3 -5,-0.2 -0.458 41.8 162.8 -68.7 118.7 22.8 22.8 11.6 112 112 A I E + 0 0 0 -7,-2.4 19,-2.8 -2,-0.4 2,-0.3 0.541 66.8 12.1-108.4 -15.2 25.0 22.2 14.6 113 113 A A E -GH 105 130A 0 -8,-1.4 -8,-1.9 17,-0.2 2,-0.3 -0.995 55.5-158.3-161.3 156.7 26.5 25.7 14.8 114 114 A G E - 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