==== Secondary Structure Definition by the program DSSP, updated CMBI version by ElmK / April 1,2000 ==== DATE=28-SEP-2012 . REFERENCE W. KABSCH AND C.SANDER, BIOPOLYMERS 22 (1983) 2577-2637 . HEADER LYASE/LYASE INHIBITOR 21-FEB-12 4DU8 . COMPND 2 MOLECULE: MEVALONATE DIPHOSPHATE DECARBOXYLASE; . SOURCE 2 ORGANISM_SCIENTIFIC: STAPHYLOCOCCUS EPIDERMIDIS; . AUTHOR M.L.BARTA,W.J.MCWHORTER,B.V.GEISBRECHT . 652 5 0 0 0 TOTAL NUMBER OF RESIDUES, NUMBER OF CHAINS, NUMBER OF SS-BRIDGES(TOTAL,INTRACHAIN,INTERCHAIN) . 27285.0 ACCESSIBLE SURFACE OF PROTEIN (ANGSTROM**2) . 492 75.5 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(J) , SAME NUMBER PER 100 RESIDUES . 10 1.5 TOTAL NUMBER OF HYDROGEN BONDS IN PARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 133 20.4 TOTAL NUMBER OF HYDROGEN BONDS IN ANTIPARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 2 0.3 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-5), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-4), SAME NUMBER PER 100 RESIDUES . 2 0.3 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-3), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-2), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-1), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+0), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+1), SAME NUMBER PER 100 RESIDUES . 51 7.8 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+2), SAME NUMBER PER 100 RESIDUES . 61 9.4 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+3), SAME NUMBER PER 100 RESIDUES . 216 33.1 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+4), SAME NUMBER PER 100 RESIDUES . 11 1.7 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+5), SAME NUMBER PER 100 RESIDUES . 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 *** HISTOGRAMS OF *** . 0 0 0 1 0 0 0 0 2 1 2 1 0 0 4 0 0 2 0 4 0 0 0 0 0 0 0 0 0 0 RESIDUES PER ALPHA HELIX . 0 0 0 2 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PARALLEL BRIDGES PER LADDER . 3 6 0 4 4 2 0 1 1 0 0 2 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 ANTIPARALLEL BRIDGES PER LADDER . 2 0 2 0 0 0 2 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 LADDERS PER SHEET . # RESIDUE AA STRUCTURE BP1 BP2 ACC N-H-->O O-->H-N N-H-->O O-->H-N TCO KAPPA ALPHA PHI PSI X-CA Y-CA Z-CA 1 -2 A T 0 0 145 0, 0.0 5,-0.0 0, 0.0 6,-0.0 0.000 360.0 360.0 360.0 168.8 49.4 12.7 30.5 2 -1 A G > - 0 0 39 1,-0.1 3,-0.8 2,-0.0 0, 0.0 -0.189 360.0-102.9 -72.1 164.7 50.5 9.0 30.2 3 0 A S T 3 S+ 0 0 122 1,-0.2 -1,-0.1 3,-0.0 0, 0.0 0.938 119.8 26.2 -50.6 -60.4 52.5 7.4 27.4 4 1 A M T 3 S+ 0 0 125 2,-0.0 2,-0.4 0, 0.0 -1,-0.2 0.322 98.5 117.1 -90.8 9.3 49.6 5.7 25.6 5 2 A V < - 0 0 21 -3,-0.8 2,-0.5 48,-0.1 48,-0.2 -0.631 46.8-163.6 -86.9 133.1 47.0 8.1 26.9 6 3 A K E +A 52 0A 109 46,-2.4 46,-3.2 -2,-0.4 2,-0.3 -0.953 15.0 175.4-118.0 114.7 45.2 10.3 24.3 7 4 A S E +A 51 0A 41 -2,-0.5 2,-0.3 44,-0.2 44,-0.2 -0.890 10.2 177.6-125.0 155.5 43.3 13.3 25.6 8 5 A G E -A 50 0A 5 42,-2.2 42,-2.7 -2,-0.3 2,-0.4 -0.996 7.7-164.6-150.7 144.3 41.4 16.2 24.1 9 6 A K E +A 49 0A 45 320,-0.4 320,-2.0 -2,-0.3 2,-0.3 -0.996 10.9 178.7-136.1 136.2 39.4 19.2 25.4 10 7 A A E -AB 48 328A 0 38,-2.1 38,-2.6 -2,-0.4 2,-0.5 -0.979 20.6-148.1-138.4 145.9 37.0 21.5 23.6 11 8 A R E +AB 47 327A 62 316,-2.9 316,-2.7 -2,-0.3 2,-0.4 -0.967 22.4 178.1-110.6 125.7 34.8 24.4 24.3 12 9 A A E -A 46 0A 0 34,-2.4 34,-3.0 -2,-0.5 2,-0.2 -0.976 18.1-137.1-128.4 142.1 31.5 24.7 22.4 13 10 A H E -A 45 0A 12 135,-0.5 32,-0.3 -2,-0.4 29,-0.1 -0.610 21.2-109.5 -99.7 155.9 28.8 27.4 22.7 14 11 A T E - 0 0 6 30,-2.3 27,-2.4 27,-0.3 2,-0.3 -0.385 30.4-153.7 -76.3 161.6 25.1 27.0 22.8 15 12 A N E -A 40 0A 10 25,-0.3 2,-0.5 -2,-0.1 25,-0.3 -0.983 6.8-140.4-135.8 147.2 23.0 28.1 19.9 16 13 A I E -A 39 0A 20 23,-2.5 23,-2.0 -2,-0.3 2,-0.4 -0.942 22.4-125.5-108.5 130.0 19.4 29.2 19.7 17 14 A A E +A 38 0A 4 -2,-0.5 21,-0.3 637,-0.3 3,-0.2 -0.587 24.8 179.6 -74.1 125.2 17.2 28.2 16.7 18 15 A L E S+ 0 0 0 19,-3.7 2,-0.6 -2,-0.4 225,-0.3 0.817 86.6 37.2 -91.5 -43.0 15.6 31.1 14.8 19 16 A I E S-A 37 0A 0 18,-2.0 18,-1.8 223,-0.1 -1,-0.4 -0.975 108.3-121.1-104.3 121.5 14.0 28.8 12.3 20 17 A K - 0 0 1 -2,-0.6 2,-0.5 16,-0.2 18,-0.1 -0.301 13.8-147.9 -76.5 138.9 13.0 25.8 14.4 21 18 A Y + 0 0 0 -2,-0.1 2,-0.2 15,-0.1 3,-0.1 -0.893 36.6 159.1 -85.0 128.8 14.0 22.2 14.0 22 19 A W + 0 0 0 -2,-0.5 174,-0.0 1,-0.4 -2,-0.0 -0.700 43.8 18.7-161.1 103.8 10.9 20.4 15.3 23 20 A G S S- 0 0 0 -2,-0.2 10,-3.0 10,-0.1 11,-0.4 0.202 70.2-119.1 103.7 134.2 10.1 16.8 14.4 24 21 A K E -H 32 0B 12 8,-0.3 3,-0.2 9,-0.1 6,-0.0 -0.828 11.4-160.6-109.1 140.1 12.4 14.1 13.0 25 22 A A E S+ 0 0 64 6,-2.4 2,-0.4 -2,-0.4 -1,-0.1 0.764 96.7 26.5 -76.9 -30.4 12.3 12.0 9.8 26 23 A D E >> -H 31 0B 60 5,-2.2 5,-2.0 1,-0.1 4,-0.9 -0.910 63.6-169.0-141.5 112.3 14.5 9.5 11.5 27 24 A E T 45S+ 0 0 88 -2,-0.4 -1,-0.1 -3,-0.2 -2,-0.0 0.804 86.1 51.0 -69.0 -33.7 14.6 9.0 15.3 28 25 A T T 45S+ 0 0 107 1,-0.2 -1,-0.2 2,-0.1 -2,-0.0 0.966 122.0 27.1 -73.6 -53.0 17.6 6.7 15.4 29 26 A Y T 45S- 0 0 102 2,-0.1 -1,-0.2 112,-0.0 -2,-0.2 0.485 109.0-119.7 -86.2 -3.5 20.1 8.8 13.4 30 27 A I T <5 + 0 0 4 -4,-0.9 -3,-0.2 1,-0.2 112,-0.1 0.995 46.4 171.1 57.9 66.7 18.4 12.0 14.3 31 28 A I E < -H 26 0B 5 -5,-2.0 -6,-2.4 129,-0.1 -5,-2.2 -0.945 25.9-135.5-102.9 128.9 17.4 13.3 10.8 32 29 A P E -H 24 0B 2 0, 0.0 -8,-0.3 0, 0.0 3,-0.0 -0.457 11.2-127.4 -87.7 156.2 15.1 16.3 11.1 33 30 A M S S+ 0 0 36 -10,-3.0 2,-0.3 -2,-0.1 -9,-0.1 0.705 91.2 12.2 -71.5 -20.4 11.8 16.9 9.1 34 31 A N S S- 0 0 1 -11,-0.4 -12,-0.1 124,-0.1 -1,-0.1 -0.973 85.7 -89.2-151.6 162.4 13.1 20.3 8.0 35 32 A N - 0 0 10 -2,-0.3 2,-0.3 -14,-0.1 122,-0.2 -0.368 40.1-155.0 -68.0 154.4 16.2 22.6 7.9 36 33 A S E - C 0 156A 0 120,-1.2 120,-2.1 -2,-0.1 2,-0.4 -0.914 7.7-152.4-133.1 160.2 16.9 24.9 10.8 37 34 A L E +AC 19 155A 0 -18,-1.8 -19,-3.7 -2,-0.3 -18,-2.0 -1.000 23.0 164.8-136.7 133.1 18.7 28.2 11.3 38 35 A S E -AC 17 154A 0 116,-2.5 116,-2.2 -2,-0.4 2,-0.4 -0.862 30.3-130.6-143.1 168.8 20.4 29.4 14.5 39 36 A V E -AC 16 153A 1 -23,-2.0 -23,-2.5 -2,-0.3 2,-0.5 -0.990 26.1-127.4-124.8 139.1 22.8 31.8 16.1 40 37 A T E -A 15 0A 3 112,-1.8 282,-2.2 -2,-0.4 2,-0.4 -0.748 24.7-130.8 -91.5 125.5 25.6 30.7 18.5 41 38 A L E -I 321 0C 0 -27,-2.4 -27,-0.3 -2,-0.5 3,-0.3 -0.590 7.6-153.9 -83.6 129.1 25.7 32.5 21.8 42 39 A D E S+ 0 0 71 278,-2.4 279,-0.1 -2,-0.4 -1,-0.1 0.863 90.6 52.1 -59.3 -39.0 28.9 34.1 23.2 43 40 A R E S+ 0 0 105 277,-0.6 2,-0.4 1,-0.1 -1,-0.2 0.775 101.5 57.6 -80.0 -23.8 27.9 33.8 26.9 44 41 A F E + 0 0 10 -3,-0.3 -30,-2.3 -30,-0.2 2,-0.3 -0.903 63.0 133.6-122.2 130.3 26.9 30.2 27.3 45 42 A Y E -AD 13 100A 53 55,-1.7 55,-3.1 -2,-0.4 2,-0.4 -0.954 40.3-120.4-162.3 174.2 29.0 27.1 26.6 46 43 A T E -AD 12 99A 0 -34,-3.0 -34,-2.4 -2,-0.3 2,-0.5 -0.970 13.5-154.5-129.3 141.5 30.2 23.7 27.6 47 44 A E E -AD 11 98A 17 51,-2.7 51,-2.7 -2,-0.4 2,-0.5 -0.980 18.1-177.4-116.9 123.9 33.8 22.5 28.2 48 45 A T E -AD 10 97A 3 -38,-2.6 -38,-2.1 -2,-0.5 2,-0.4 -0.969 10.1-171.0-132.2 121.7 34.4 18.8 27.7 49 46 A K E -AD 9 96A 65 47,-3.1 47,-2.7 -2,-0.5 2,-0.4 -0.882 6.0-168.8-105.1 138.2 37.6 16.9 28.2 50 47 A V E -AD 8 95A 0 -42,-2.7 -42,-2.2 -2,-0.4 2,-0.5 -0.981 3.3-165.7-125.3 144.0 38.0 13.2 27.2 51 48 A T E -AD 7 94A 21 43,-1.9 43,-2.5 -2,-0.4 2,-0.6 -0.973 4.9-159.2-132.8 114.2 40.8 10.9 28.1 52 49 A F E +AD 6 93A 0 -46,-3.2 -46,-2.4 -2,-0.5 41,-0.2 -0.855 17.2 169.8 -95.3 125.2 41.3 7.6 26.3 53 50 A D E > - D 0 92A 27 39,-1.3 3,-2.2 -2,-0.6 39,-0.5 -0.994 35.7-138.6-135.8 130.2 43.4 5.0 28.1 54 51 A P T 3 S+ 0 0 54 0, 0.0 -1,-0.1 0, 0.0 -49,-0.0 0.770 105.4 58.8 -54.2 -30.7 43.8 1.4 27.1 55 52 A D T 3 S+ 0 0 134 -3,-0.0 2,-0.2 37,-0.0 -3,-0.0 0.586 81.3 106.7 -79.5 -10.9 43.5 0.3 30.8 56 53 A F < - 0 0 75 -3,-2.2 36,-0.4 36,-0.2 3,-0.1 -0.515 53.0-166.3 -68.6 136.8 40.1 1.9 31.0 57 54 A T + 0 0 122 1,-0.2 2,-0.3 -2,-0.2 -1,-0.1 0.820 66.0 1.4 -98.1 -38.6 37.3 -0.7 31.1 58 55 A E S S- 0 0 93 34,-0.1 2,-0.3 35,-0.0 -1,-0.2 -0.874 87.1 -85.3-139.0 170.7 34.2 1.3 30.4 59 56 A D - 0 0 1 -2,-0.3 2,-0.4 33,-0.1 35,-0.2 -0.614 38.9-160.0 -79.9 144.0 33.4 5.0 29.7 60 57 A C E -e 94 0A 40 33,-2.3 35,-3.9 -2,-0.3 2,-0.5 -0.991 2.5-159.8-127.7 130.1 33.0 7.3 32.7 61 58 A L E -e 95 0A 18 -2,-0.4 7,-0.9 33,-0.2 2,-0.5 -0.946 6.5-174.9-114.5 118.8 31.2 10.6 32.6 62 59 A I E -eF 96 67A 16 33,-3.7 35,-3.1 -2,-0.5 2,-0.6 -0.958 8.2-167.0-113.0 120.8 31.9 13.2 35.3 63 60 A L E > S-eF 97 66A 37 3,-3.0 3,-1.8 -2,-0.5 35,-0.2 -0.926 70.0 -20.6-116.9 117.0 29.7 16.3 35.0 64 61 A N T 3 S- 0 0 57 33,-3.2 -1,-0.2 -2,-0.6 34,-0.1 0.925 129.4 -50.2 50.3 48.0 30.5 19.4 37.0 65 62 A G T 3 S+ 0 0 59 32,-0.3 2,-0.5 -3,-0.2 -1,-0.3 0.439 121.0 109.2 75.4 0.1 32.6 17.3 39.4 66 63 A N E < -F 63 0A 50 -3,-1.8 -3,-3.0 575,-0.0 -1,-0.2 -0.922 67.8-125.1-120.0 131.2 29.7 14.8 39.8 67 64 A E E -F 62 0A 139 -2,-0.5 2,-0.2 -5,-0.2 -5,-0.2 -0.288 31.7-130.3 -58.4 149.6 29.2 11.2 38.6 68 65 A V - 0 0 34 -7,-0.9 -7,-0.1 1,-0.2 -1,-0.1 -0.605 27.2 -87.3-102.4 164.9 26.0 10.7 36.6 69 66 A N > - 0 0 29 -2,-0.2 4,-2.3 1,-0.1 -1,-0.2 -0.061 40.8-102.0 -65.1 170.1 23.3 8.0 37.1 70 67 A A H > S+ 0 0 80 2,-0.2 4,-1.8 1,-0.2 -1,-0.1 0.777 123.7 56.0 -66.2 -24.4 23.5 4.5 35.5 71 68 A K H > S+ 0 0 153 2,-0.2 4,-2.2 1,-0.2 3,-0.4 0.988 110.1 40.7 -66.6 -62.9 21.0 5.8 33.0 72 69 A E H > S+ 0 0 43 1,-0.2 4,-1.8 2,-0.2 -2,-0.2 0.828 115.2 55.8 -53.9 -33.7 23.1 8.8 31.8 73 70 A K H X S+ 0 0 66 -4,-2.3 4,-1.6 2,-0.2 -1,-0.2 0.889 105.6 49.0 -68.6 -40.4 26.1 6.4 32.0 74 71 A E H X S+ 0 0 101 -4,-1.8 4,-1.8 -3,-0.4 -2,-0.2 0.906 110.8 52.1 -65.2 -40.8 24.5 3.9 29.6 75 72 A K H X S+ 0 0 88 -4,-2.2 4,-2.0 1,-0.2 -2,-0.2 0.870 108.7 48.1 -63.9 -40.8 23.6 6.7 27.2 76 73 A I H X S+ 0 0 10 -4,-1.8 4,-2.4 2,-0.2 -1,-0.2 0.846 108.9 55.2 -72.0 -30.9 27.2 8.1 27.0 77 74 A Q H X S+ 0 0 28 -4,-1.6 4,-1.3 2,-0.2 -2,-0.2 0.886 107.9 48.6 -68.0 -37.6 28.6 4.6 26.5 78 75 A N H X S+ 0 0 86 -4,-1.8 4,-1.1 2,-0.2 3,-0.3 0.933 111.9 49.0 -65.8 -43.9 26.3 4.1 23.5 79 76 A Y H >X S+ 0 0 10 -4,-2.0 4,-1.4 1,-0.2 3,-0.7 0.914 108.6 53.3 -60.8 -42.0 27.4 7.5 22.1 80 77 A M H 3X S+ 0 0 0 -4,-2.4 4,-2.9 1,-0.3 -1,-0.2 0.771 97.8 65.4 -68.3 -23.7 31.0 6.6 22.6 81 78 A N H 3X S+ 0 0 70 -4,-1.3 4,-2.4 -3,-0.3 -1,-0.3 0.876 99.3 53.0 -60.6 -37.3 30.5 3.4 20.7 82 79 A I H S+ 0 0 10 -4,-2.9 5,-2.0 2,-0.2 4,-1.5 0.910 111.4 46.7 -55.4 -46.8 35.0 4.0 18.7 85 82 A D H <5S+ 0 0 136 -4,-2.4 3,-0.5 2,-0.2 -1,-0.2 0.958 112.6 50.2 -60.1 -50.4 33.7 2.7 15.4 86 83 A L H <5S+ 0 0 83 -4,-2.4 -2,-0.2 1,-0.2 -1,-0.2 0.898 116.6 40.9 -52.7 -45.8 34.5 5.9 13.7 87 84 A A H <5S- 0 0 30 -4,-2.7 -1,-0.2 -5,-0.2 -2,-0.2 0.664 106.4-127.3 -78.7 -17.5 38.1 5.8 15.1 88 85 A G T <5S+ 0 0 72 -4,-1.5 2,-0.2 -3,-0.5 -3,-0.2 0.825 72.0 105.6 71.2 33.3 38.5 2.1 14.5 89 86 A N < - 0 0 38 -5,-2.0 -1,-0.3 -6,-0.2 -2,-0.1 -0.768 67.4-138.8-134.0-178.6 39.5 1.4 18.1 90 87 A R + 0 0 224 -2,-0.2 2,-0.2 -3,-0.1 -6,-0.1 0.041 52.0 138.0-131.9 21.9 38.1 0.0 21.3 91 88 A L - 0 0 42 -7,-0.1 2,-0.3 -39,-0.0 -2,-0.1 -0.507 42.3-145.6 -72.4 136.2 39.5 2.5 23.8 92 89 A H E -D 53 0A 48 -39,-0.5 -39,-1.3 -36,-0.4 2,-0.3 -0.698 18.2-141.6 -97.0 158.6 37.1 3.7 26.5 93 90 A A E -D 52 0A 0 -2,-0.3 -33,-2.3 -41,-0.2 2,-0.6 -0.875 15.8-137.0-126.9 150.0 37.2 7.2 27.8 94 91 A R E -De 51 60A 96 -43,-2.5 -43,-1.9 -2,-0.3 2,-0.6 -0.943 29.8-160.9 -96.0 122.9 37.0 9.2 31.0 95 92 A I E -De 50 61A 2 -35,-3.9 -33,-3.7 -2,-0.6 2,-0.6 -0.921 11.0-174.9-111.5 117.9 34.8 12.3 30.1 96 93 A E E +De 49 62A 64 -47,-2.7 -47,-3.1 -2,-0.6 2,-0.4 -0.961 14.1 177.3-112.4 115.2 35.0 15.3 32.4 97 94 A S E -De 48 63A 6 -35,-3.1 -33,-3.2 -2,-0.6 2,-0.4 -0.936 17.7-167.0-123.5 143.2 32.5 18.0 31.4 98 95 A E E -D 47 0A 50 -51,-2.7 -51,-2.7 -2,-0.4 2,-0.6 -0.998 14.0-147.0-125.6 127.2 31.5 21.4 32.8 99 96 A N E -D 46 0A 28 -2,-0.4 -53,-0.2 -53,-0.2 2,-0.2 -0.854 14.3-176.2 -93.4 120.6 28.4 23.2 31.7 100 97 A Y E +D 45 0A 109 -55,-3.1 -55,-1.7 -2,-0.6 -54,-0.1 -0.492 55.0 109.6-105.6 57.8 28.6 27.0 31.7 101 98 A V S S- 0 0 12 -2,-0.2 2,-2.6 -57,-0.2 3,-0.3 -0.991 81.0-115.8-135.3 133.2 24.9 27.2 30.7 102 99 A P > + 0 0 5 0, 0.0 5,-2.2 0, 0.0 3,-0.2 -0.421 53.3 154.3 -69.9 70.3 22.0 28.3 32.9 103 100 A T T > 5 + 0 0 44 -2,-2.6 3,-2.1 1,-0.2 534,-0.1 0.859 67.7 57.8 -71.3 -39.1 20.4 24.9 32.9 104 101 A A T 3 5S+ 0 0 4 -3,-0.3 -1,-0.2 1,-0.3 328,-0.1 0.844 104.8 54.3 -55.0 -34.3 18.6 25.4 36.2 105 102 A A T 3 5S- 0 0 7 -3,-0.2 -1,-0.3 326,-0.1 81,-0.2 0.344 130.1 -98.1 -84.9 5.2 17.0 28.5 34.5 106 103 A G T < 5S+ 0 0 42 -3,-2.1 2,-0.3 1,-0.3 -3,-0.2 0.477 75.2 141.9 95.7 2.8 15.8 26.2 31.7 107 104 A L < - 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