==== Secondary Structure Definition by the program DSSP, updated CMBI version by ElmK / April 1,2000 ==== DATE=11-JAN-2013 . REFERENCE W. KABSCH AND C.SANDER, BIOPOLYMERS 22 (1983) 2577-2637 . HEADER HYDROLASE/IMMUNE SYSTEM 24-FEB-12 4DW2 . COMPND 2 MOLECULE: UROKINASE-TYPE PLASMINOGEN ACTIVATOR; . SOURCE 2 ORGANISM_SCIENTIFIC: HOMO SAPIENS; . AUTHOR L.JIANG,K.A.BOTKJAER,L.M.ANDERSEN,C.YUAN,P.A.ANDREASEN,M.HUA . 645 9 8 6 2 TOTAL NUMBER OF RESIDUES, NUMBER OF CHAINS, NUMBER OF SS-BRIDGES(TOTAL,INTRACHAIN,INTERCHAIN) . 26260.0 ACCESSIBLE SURFACE OF PROTEIN (ANGSTROM**2) . 436 67.6 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(J) , SAME NUMBER PER 100 RESIDUES . 10 1.6 TOTAL NUMBER OF HYDROGEN BONDS IN PARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 230 35.7 TOTAL NUMBER OF HYDROGEN BONDS IN ANTIPARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 4 0.6 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-5), SAME NUMBER PER 100 RESIDUES . 8 1.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-4), SAME NUMBER PER 100 RESIDUES . 7 1.1 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-3), SAME NUMBER PER 100 RESIDUES . 1 0.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-2), SAME NUMBER PER 100 RESIDUES . 2 0.3 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-1), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+0), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+1), SAME NUMBER PER 100 RESIDUES . 100 15.5 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+2), SAME NUMBER PER 100 RESIDUES . 44 6.8 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+3), SAME NUMBER PER 100 RESIDUES . 25 3.9 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+4), SAME NUMBER PER 100 RESIDUES . 3 0.5 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+5), SAME NUMBER PER 100 RESIDUES . 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 *** HISTOGRAMS OF *** . 0 0 0 0 1 1 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 RESIDUES PER ALPHA HELIX . 3 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PARALLEL BRIDGES PER LADDER . 9 6 6 9 6 8 2 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 ANTIPARALLEL BRIDGES PER LADDER . 1 0 4 2 1 1 2 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 LADDERS PER SHEET . # RESIDUE AA STRUCTURE BP1 BP2 ACC N-H-->O O-->H-N N-H-->O O-->H-N TCO KAPPA ALPHA PHI PSI X-CA Y-CA Z-CA 1 21 U F 0 0 164 0, 0.0 2,-0.9 0, 0.0 491,-0.1 0.000 360.0 360.0 360.0 -1.9 -12.9 -1.5 -8.4 2 22 U T > + 0 0 67 1,-0.1 3,-0.8 142,-0.1 4,-0.1 -0.835 360.0 178.9 -98.3 99.9 -13.8 0.7 -11.4 3 23 U T T 3 + 0 0 7 -2,-0.9 3,-0.3 142,-0.3 -1,-0.1 0.206 55.8 92.7 -94.1 5.0 -17.6 0.7 -11.1 4 24 U I T 3 S+ 0 0 0 1,-0.2 -1,-0.2 4,-0.1 6,-0.2 0.671 73.6 73.4 -75.4 -16.3 -18.7 2.8 -14.0 5 25 U E S < S+ 0 0 87 -3,-0.8 2,-0.6 2,-0.1 -1,-0.2 0.913 95.6 56.7 -62.1 -40.5 -19.0 -0.5 -15.9 6 26 U N S S+ 0 0 58 -3,-0.3 141,-0.0 1,-0.3 100,-0.0 -0.827 128.4 8.4 -76.4 127.3 -22.0 -0.6 -13.7 7 27 U Q S > S+ 0 0 11 -2,-0.6 3,-1.2 1,-0.1 -1,-0.3 0.960 77.7 170.6 62.8 57.7 -23.7 2.7 -14.7 8 28 U P T 3 + 0 0 9 0, 0.0 101,-4.5 0, 0.0 45,-0.2 0.015 63.0 76.7 -93.4 31.3 -21.5 3.6 -17.7 9 29 U W T 3 + 0 0 23 99,-0.2 17,-2.4 101,-0.1 102,-0.2 0.481 68.5 120.7 -98.9 -12.0 -23.7 6.4 -18.9 10 30 U F B < -A 25 0A 1 -3,-1.2 2,-0.6 15,-0.2 43,-0.5 -0.283 50.2-160.8 -64.4 128.1 -22.3 8.3 -16.1 11 31 U A - 0 0 0 13,-2.1 2,-1.1 22,-0.2 41,-0.2 -0.943 9.5-151.0-110.7 120.3 -20.5 11.4 -17.4 12 32 U A E -B 51 0A 2 39,-1.6 39,-2.7 -2,-0.6 2,-0.6 -0.786 17.2-162.0 -88.7 102.0 -18.0 13.1 -15.2 13 33 U I E -BC 50 22A 0 9,-2.4 8,-2.1 -2,-1.1 9,-1.7 -0.756 12.3-177.6 -89.0 121.6 -18.3 16.7 -16.3 14 34 U Y E -BC 49 20A 0 35,-3.6 35,-2.3 -2,-0.6 2,-0.5 -0.660 24.8-128.8-112.0 169.3 -15.3 18.6 -15.2 15 35 U R E BC 48 19A 56 4,-4.2 4,-3.4 33,-0.2 33,-0.3 -0.986 360.0 360.0-119.8 125.9 -14.5 22.2 -15.6 16 36 U R 0 0 117 31,-2.6 -1,-0.2 -2,-0.5 32,-0.1 0.976 360.0 360.0 -77.6 360.0 -11.1 22.9 -17.0 17 ! 0 0 0 0, 0.0 0, 0.0 0, 0.0 0, 0.0 0.000 360.0 360.0 360.0 360.0 0.0 0.0 0.0 18 38 U V 0 0 86 0, 0.0 2,-0.4 0, 0.0 -2,-0.3 0.000 360.0 360.0 360.0 122.4 -9.9 20.9 -13.9 19 39 U T E -C 15 0A 77 -4,-3.4 -4,-4.2 -3,-0.0 2,-0.6 -0.927 360.0-110.9-120.1 152.6 -12.5 20.8 -11.2 20 40 U Y E +C 14 0A 41 -2,-0.4 -6,-0.2 -6,-0.3 3,-0.1 -0.657 40.3 167.6 -85.2 120.3 -15.4 18.4 -11.0 21 41 U V E - 0 0 39 -8,-2.1 2,-0.3 -2,-0.6 -7,-0.2 0.884 53.9 -20.7 -96.1 -49.5 -18.6 20.2 -11.5 22 42 U a E -C 13 0A 2 -9,-1.7 -9,-2.4 161,-0.1 -1,-0.4 -0.968 53.7-110.3-159.6 158.2 -21.3 17.7 -12.1 23 43 U G - 0 0 0 159,-2.6 12,-0.5 -2,-0.3 2,-0.3 -0.298 34.6-176.5 -84.7-176.9 -22.2 14.3 -13.1 24 44 U G - 0 0 0 -13,-0.2 -13,-2.1 10,-0.2 2,-0.4 -0.901 19.8-125.4-167.5 177.6 -24.0 13.6 -16.3 25 45 U S E -AD 10 33A 0 8,-2.5 8,-3.2 -15,-0.3 2,-0.8 -0.985 21.2-122.3-145.6 131.3 -25.5 10.6 -18.1 26 46 U L E + D 0 32A 2 -17,-2.4 85,-4.0 83,-0.4 6,-0.2 -0.663 38.5 163.8 -74.8 109.2 -24.7 9.5 -21.7 27 47 U I S S+ 0 0 15 4,-2.2 -1,-0.2 -2,-0.8 5,-0.2 0.812 73.7 3.5 -90.7 -39.5 -28.1 9.5 -23.4 28 48 U S S S- 0 0 26 3,-2.0 -1,-0.4 -3,-0.2 74,-0.2 -0.902 81.8-105.6-137.4 161.8 -26.5 9.4 -26.8 29 49 U P S S+ 0 0 54 0, 0.0 73,-2.3 0, 0.0 3,-0.1 0.857 125.3 37.8 -58.4 -33.1 -22.9 9.0 -28.0 30 50 U b S S+ 0 0 35 71,-0.2 64,-6.7 1,-0.1 2,-0.6 0.847 115.3 60.3 -80.6 -38.7 -23.0 12.7 -28.9 31 51 U W E - E 0 93A 22 62,-0.3 -4,-2.2 193,-0.1 -3,-2.0 -0.817 54.5-169.1-114.0 115.2 -25.1 13.9 -26.0 32 52 U V E -DE 26 92A 0 60,-4.3 60,-2.5 -2,-0.6 2,-0.3 -0.860 20.0-146.4 -91.9 124.2 -24.5 13.8 -22.3 33 53 U I E +DE 25 91A 0 -8,-3.2 -8,-2.5 -2,-0.6 2,-0.3 -0.695 34.5 144.1 -85.2 145.4 -27.6 14.8 -20.3 34 54 U S E - E 0 90A 0 56,-0.6 56,-5.0 -2,-0.3 2,-0.2 -0.910 51.0 -67.8-159.8-177.8 -27.0 16.6 -17.1 35 55 U A E - E 0 89A 0 -12,-0.5 4,-0.4 54,-0.3 54,-0.3 -0.605 30.7-152.0 -79.2 148.2 -28.2 19.3 -14.7 36 56 U T S >> S+ 0 0 0 52,-3.2 4,-2.3 -2,-0.2 3,-0.8 0.844 87.3 75.4 -85.4 -37.3 -28.0 22.9 -16.0 37 57 U H T 34 S+ 0 0 27 51,-0.3 4,-0.5 1,-0.3 -1,-0.1 0.714 89.8 66.1 -48.2 -24.9 -27.6 24.5 -12.6 38 58 U a T >4 S+ 0 0 10 50,-0.2 3,-0.7 1,-0.1 4,-0.4 0.996 112.1 25.4 -53.2 -65.5 -24.1 23.2 -12.9 39 59 U F T <4 S+ 0 0 0 -3,-0.8 3,-0.3 -4,-0.4 -2,-0.2 0.664 96.7 85.4 -83.3 -26.5 -23.1 25.4 -15.8 40 60 U I T 3< S+ 0 0 48 -4,-2.3 -1,-0.3 1,-0.3 -3,-0.1 0.887 88.0 60.4 -47.2 -33.3 -25.3 28.3 -15.5 41 60AU D S < S+ 0 0 111 -3,-0.7 -1,-0.3 -4,-0.5 -2,-0.2 0.985 130.7 7.5 -66.3 -40.9 -22.7 29.5 -13.2 42 60BU Y S S- 0 0 119 -4,-0.4 -1,-0.3 -3,-0.3 -2,-0.1 -0.827 75.5-173.2-138.1 109.1 -20.4 29.3 -16.1 43 60CU P + 0 0 72 0, 0.0 2,-0.3 0, 0.0 29,-0.1 0.333 38.9 122.4 -85.8 2.4 -22.2 28.5 -19.3 44 61 U K > - 0 0 113 -5,-0.2 4,-0.8 1,-0.2 3,-0.2 -0.514 52.2-152.0 -78.9 131.7 -19.3 28.0 -21.7 45 62 U K T >4 S+ 0 0 65 -2,-0.3 24,-0.8 1,-0.3 3,-0.6 0.833 101.3 60.0 -60.5 -34.4 -18.9 24.8 -23.6 46 62AU E T 34 S+ 0 0 91 1,-0.2 -1,-0.3 22,-0.1 -30,-0.2 0.952 97.2 61.9 -61.1 -39.3 -15.2 25.6 -23.7 47 63 U D T 34 S+ 0 0 33 -3,-0.2 -31,-2.6 -31,-0.1 2,-0.3 0.788 96.7 63.9 -57.8 -34.9 -15.2 25.5 -20.0 48 64 U Y E << -B 15 0A 2 -4,-0.8 21,-0.8 -3,-0.6 2,-0.4 -0.640 55.0-163.1-109.1 146.7 -16.2 22.0 -19.5 49 65 U I E -BF 14 68A 2 -35,-2.3 -35,-3.6 -2,-0.3 2,-0.6 -0.992 15.1-148.8-122.1 128.7 -14.8 18.7 -20.2 50 66 U V E -BF 13 67A 0 17,-3.9 17,-2.8 -2,-0.4 2,-0.3 -0.886 11.6-163.1-100.6 126.1 -16.9 15.6 -20.2 51 67 U Y E -BF 12 66A 15 -39,-2.7 -39,-1.6 -2,-0.6 2,-0.3 -0.791 5.3-171.5 -97.3 147.8 -15.4 12.4 -19.2 52 68 U L E + F 0 65A 2 13,-2.4 13,-1.7 -2,-0.3 56,-0.1 -0.886 68.7 22.9-127.5 167.2 -16.9 9.1 -19.8 53 69 U G S S+ 0 0 0 -43,-0.5 2,-0.5 54,-0.4 55,-0.1 0.802 87.1 139.7 47.7 29.5 -15.9 5.6 -18.6 54 70 U R + 0 0 8 -44,-0.4 -1,-0.2 9,-0.1 6,-0.1 -0.937 31.3 179.8-104.3 121.9 -14.1 7.4 -15.8 55 71 U S + 0 0 2 -2,-0.5 90,-5.6 4,-0.1 2,-0.3 0.494 63.3 53.9 -91.9 -5.2 -14.6 5.7 -12.4 56 72 U R B S-H 144 0B 58 3,-0.4 88,-0.2 88,-0.3 5,-0.2 -0.923 71.0-132.1-147.0 147.7 -12.6 8.1 -10.4 57 73 U L S S+ 0 0 32 86,-0.7 -1,-0.2 -2,-0.3 86,-0.1 0.995 100.1 9.7 -76.9 -65.8 -12.4 11.8 -9.7 58 74 U N S S+ 0 0 123 2,-0.1 2,-0.5 -38,-0.0 -1,-0.2 0.249 116.4 80.5-101.1 14.8 -8.8 13.0 -10.1 59 75 U S S S- 0 0 35 84,-0.1 2,-1.1 0, 0.0 -3,-0.4 -0.979 76.9-135.2-120.0 124.1 -7.5 9.8 -11.6 60 76 U N - 0 0 98 -2,-0.5 2,-0.1 -5,-0.1 -3,-0.1 -0.662 26.6-156.8 -74.9 101.7 -8.0 9.1 -15.2 61 77 U T > - 0 0 24 -2,-1.1 3,-1.0 -5,-0.2 2,-0.1 -0.482 26.3 -97.6 -72.7 155.8 -9.1 5.5 -15.1 62 78 U Q T 3 S+ 0 0 191 1,-0.2 3,-0.1 -2,-0.1 -7,-0.1 -0.418 109.0 36.3 -78.2 153.9 -8.5 3.5 -18.3 63 79 U G T 3 S+ 0 0 41 1,-0.4 -1,-0.2 -2,-0.1 -9,-0.1 0.077 88.8 131.6 89.6 -23.1 -11.3 3.1 -20.7 64 80 U E < - 0 0 37 -3,-1.0 2,-0.5 -11,-0.1 -1,-0.4 -0.384 46.8-152.1 -60.8 132.1 -12.3 6.6 -19.7 65 81 U M E -F 52 0A 37 -13,-1.7 -13,-2.4 -3,-0.1 2,-0.4 -0.950 3.3-144.5-113.1 126.8 -13.0 8.8 -22.7 66 82 U K E -F 51 0A 79 -2,-0.5 31,-0.6 -15,-0.2 2,-0.4 -0.702 21.6-176.9 -85.5 137.6 -12.4 12.6 -22.5 67 83 U F E -FG 50 96A 0 -17,-2.8 -17,-3.9 -2,-0.4 29,-0.3 -0.998 22.2-141.3-137.6 138.0 -14.8 14.9 -24.4 68 84 U E E -FG 49 95A 74 27,-3.4 27,-3.9 -2,-0.4 2,-0.9 -0.545 37.8-107.4 -79.2 159.3 -15.1 18.6 -25.1 69 85 U V E - G 0 94A 5 -21,-0.8 25,-0.3 -24,-0.8 3,-0.2 -0.831 30.0-171.7 -91.6 104.0 -18.6 19.9 -25.1 70 86 U E E S+ 0 0 97 23,-1.5 2,-0.4 -2,-0.9 -1,-0.2 0.994 80.5 10.5 -56.1 -57.5 -19.4 20.7 -28.7 71 87 U N E - G 0 93A 76 22,-0.6 22,-2.5 -3,-0.2 2,-0.4 -0.984 67.1-156.8-128.4 130.4 -22.6 22.3 -27.5 72 88 U L E - G 0 92A 21 -2,-0.4 2,-0.4 -3,-0.2 20,-0.2 -0.831 15.1-172.8 -94.9 138.1 -23.7 23.3 -24.0 73 89 U I E - G 0 91A 19 18,-2.2 18,-1.3 -2,-0.4 2,-0.3 -0.927 4.2-174.2-140.1 112.9 -27.4 23.6 -23.7 74 90 U L E - G 0 90A 37 -2,-0.4 2,-0.4 16,-0.2 16,-0.2 -0.788 42.7 -92.0-104.4 147.6 -29.0 24.9 -20.5 75 91 U H > - 0 0 23 14,-1.0 3,-3.3 12,-0.3 11,-0.1 -0.458 27.4-140.3 -65.5 119.1 -32.8 24.9 -20.1 76 92 U K T 3 S+ 0 0 170 -2,-0.4 -1,-0.2 1,-0.3 11,-0.0 0.739 105.3 43.5 -51.4 -20.0 -34.0 28.2 -21.2 77 93 U D T 3 S+ 0 0 93 10,-0.1 -1,-0.3 2,-0.1 9,-0.1 0.358 78.4 136.6-107.1 1.9 -36.3 28.0 -18.3 78 94 U Y < - 0 0 36 -3,-3.3 2,-0.4 9,-0.2 9,-0.2 -0.302 31.8-173.5 -59.4 133.1 -34.0 26.7 -15.6 79 95 U S E -I 86 0C 51 7,-1.7 7,-3.5 2,-0.0 2,-0.8 -0.988 17.2-147.7-130.4 130.8 -34.6 28.6 -12.5 80 96 U A E -I 85 0C 58 -2,-0.4 5,-0.2 5,-0.2 2,-0.1 -0.874 16.8-165.2-102.1 116.3 -32.6 28.4 -9.4 81 97 U D - 0 0 77 3,-3.9 5,-0.0 -2,-0.8 -2,-0.0 -0.379 42.8-100.9 -83.2 172.7 -34.6 28.9 -6.4 82 97AU T S S+ 0 0 148 -2,-0.1 -1,-0.1 3,-0.0 3,-0.1 0.879 122.9 8.4 -56.6 -33.5 -33.0 29.5 -3.1 83 97BU L S S+ 0 0 92 1,-0.1 -2,-0.1 119,-0.0 81,-0.0 0.743 124.2 45.1-109.2 -84.5 -33.9 25.9 -2.8 84 98 U A - 0 0 2 -4,-0.1 -3,-3.9 1,-0.1 2,-0.5 -0.140 64.0-136.0 -70.5 161.1 -35.2 24.0 -5.8 85 99 U H E -I 80 0C 36 -5,-0.2 3,-0.5 124,-0.1 2,-0.4 -0.976 18.5-146.5-115.4 131.9 -33.9 24.1 -9.3 86 100 U H E S-I 79 0C 40 -7,-3.5 -7,-1.7 -2,-0.5 -11,-0.1 -0.823 76.3 -5.5-106.5 138.7 -36.4 24.4 -12.1 87 101 U N S S+ 0 0 36 -2,-0.4 -12,-0.3 -9,-0.2 -1,-0.3 0.911 78.2 171.2 48.4 47.7 -36.0 22.8 -15.4 88 102 U D + 0 0 0 -3,-0.5 -52,-3.2 -52,-0.1 -51,-0.3 0.346 28.9 126.8 -75.4 11.1 -32.5 21.8 -14.2 89 103 U I E -E 35 0A 0 -54,-0.3 -14,-1.0 -53,-0.1 -54,-0.3 -0.483 37.4-176.9 -65.3 126.5 -32.1 19.6 -17.3 90 104 U A E -EG 34 74A 0 -56,-5.0 -56,-0.6 -2,-0.2 2,-0.3 -0.587 11.1-148.5-108.5-177.0 -28.9 20.5 -19.1 91 105 U L E -EG 33 73A 0 -18,-1.3 -18,-2.2 -2,-0.2 2,-0.5 -0.924 7.9-168.2-155.1 135.4 -27.6 19.0 -22.4 92 106 U L E -EG 32 72A 0 -60,-2.5 -60,-4.3 -2,-0.3 2,-0.5 -0.923 12.2-157.5-132.2 109.7 -24.1 18.4 -23.7 93 107 U K E -EG 31 71A 52 -22,-2.5 -23,-1.5 -2,-0.5 -22,-0.6 -0.701 31.7-150.4 -71.2 126.2 -23.2 17.5 -27.2 94 108 U I E - 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