==== Secondary Structure Definition by the program DSSP, updated CMBI version by ElmK / April 1,2000 ==== DATE=30-DEC-2009 . REFERENCE W. KABSCH AND C.SANDER, BIOPOLYMERS 22 (1983) 2577-2637 . HEADER GENE REGULATION 25-NOV-05 2F5G . COMPND 2 MOLECULE: TRANSPOSASE, PUTATIVE; . SOURCE 2 ORGANISM_SCIENTIFIC: SULFOLOBUS SOLFATARICUS; . AUTHOR H.H.LEE,J.Y.YOON,H.S.KIM,J.Y.KANG,K.H.KIM,D.J.KIM,S.W.SUH . 260 2 0 0 0 TOTAL NUMBER OF RESIDUES, NUMBER OF CHAINS, NUMBER OF SS-BRIDGES(TOTAL,INTRACHAIN,INTERCHAIN) . 13098.0 ACCESSIBLE SURFACE OF PROTEIN (ANGSTROM**2) . 197 75.8 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(J) , SAME NUMBER PER 100 RESIDUES . 10 3.8 TOTAL NUMBER OF HYDROGEN BONDS IN PARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 58 22.3 TOTAL NUMBER OF HYDROGEN BONDS IN ANTIPARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-5), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-4), SAME NUMBER PER 100 RESIDUES . 4 1.5 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-3), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-2), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-1), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+0), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+1), SAME NUMBER PER 100 RESIDUES . 19 7.3 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+2), SAME NUMBER PER 100 RESIDUES . 15 5.8 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+3), SAME NUMBER PER 100 RESIDUES . 91 35.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+4), SAME NUMBER PER 100 RESIDUES . 2 0.8 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+5), SAME NUMBER PER 100 RESIDUES . 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 *** HISTOGRAMS OF *** . 0 0 0 0 0 0 0 0 0 0 0 0 0 0 2 0 1 1 2 0 0 0 0 0 0 0 0 0 0 0 RESIDUES PER ALPHA HELIX . 0 0 0 2 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PARALLEL BRIDGES PER LADDER . 3 0 2 0 2 2 2 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 ANTIPARALLEL BRIDGES PER LADDER . 0 0 0 0 2 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 LADDERS PER SHEET . # RESIDUE AA STRUCTURE BP1 BP2 ACC N-H-->O O-->H-N N-H-->O O-->H-N TCO KAPPA ALPHA PHI PSI X-CA Y-CA Z-CA 1 2 A E 0 0 199 0, 0.0 2,-0.1 0, 0.0 9,-0.1 0.000 360.0 360.0 360.0 164.8 68.4 16.2 24.6 2 3 A L - 0 0 73 9,-0.1 2,-0.2 7,-0.0 9,-0.2 -0.383 360.0-163.6 -64.2 141.0 67.1 19.8 24.7 3 4 A K E -A 10 0A 55 7,-2.9 7,-3.2 -2,-0.1 2,-0.4 -0.733 6.5-140.5-122.1 172.2 69.7 22.3 23.5 4 5 A S E -A 9 0A 77 5,-0.3 5,-0.2 -2,-0.2 2,-0.1 -0.998 13.3-177.0-142.5 139.5 70.0 26.1 23.8 5 6 A T E > -A 8 0A 51 3,-1.9 3,-0.6 -2,-0.4 5,-0.0 -0.341 58.4 -57.1-112.5-164.7 71.2 28.9 21.6 6 7 A R T 3 S- 0 0 194 1,-0.2 228,-0.1 -2,-0.1 -2,-0.0 0.925 132.0 -3.9 -43.0 -64.6 71.6 32.7 22.1 7 8 A H T 3 S+ 0 0 54 226,-0.1 228,-1.3 227,-0.0 2,-0.3 -0.126 121.1 77.4-125.9 41.0 68.0 33.2 23.1 8 9 A T E < -Ab 5 235A 4 -3,-0.6 -3,-1.9 226,-0.2 2,-0.4 -0.986 52.7-152.7-149.2 159.1 66.3 29.8 22.7 9 10 A K E +Ab 4 236A 49 226,-1.4 228,-2.1 -2,-0.3 2,-0.3 -0.992 40.8 172.7-123.4 129.7 65.8 26.4 24.2 10 11 A Y E -Ab 3 237A 14 -7,-3.2 -7,-2.9 -2,-0.4 2,-0.3 -0.985 41.1-172.2-151.1 159.5 65.1 23.8 21.5 11 12 A L E + b 0 238A 26 226,-1.8 228,-3.0 -2,-0.3 2,-0.5 -0.893 24.0 170.0-144.7 101.9 64.5 20.3 20.3 12 13 A C + 0 0 0 -2,-0.3 54,-3.0 54,-0.2 2,-0.4 -0.840 3.3 166.9-125.0 95.8 64.3 20.2 16.5 13 14 A N E -FG 65 109B 23 96,-0.7 96,-2.1 -2,-0.5 2,-0.4 -0.900 8.2-178.2-109.3 137.6 64.3 16.7 15.1 14 15 A Y E -FG 64 108B 0 50,-2.5 50,-2.5 -2,-0.4 2,-0.5 -0.991 22.0-144.3-137.1 145.4 63.4 15.9 11.5 15 16 A H E -FG 63 107B 15 92,-2.8 92,-1.8 -2,-0.4 2,-0.4 -0.919 21.6-171.7-103.3 127.1 63.1 12.8 9.3 16 17 A F E +FG 62 106B 0 46,-2.9 46,-2.9 -2,-0.5 2,-0.4 -0.974 4.9 179.2-120.2 137.8 64.2 13.4 5.7 17 18 A V E +FG 61 105B 0 88,-2.0 88,-2.7 -2,-0.4 2,-0.3 -0.997 10.3 151.3-140.2 132.2 63.7 10.7 3.0 18 19 A W E -F 60 0B 0 42,-1.6 42,-2.6 -2,-0.4 86,-0.1 -0.974 26.8-134.1-153.0 163.7 64.5 11.0 -0.7 19 20 A I - 0 0 1 84,-0.3 82,-1.6 -2,-0.3 40,-0.2 -0.852 24.5-103.2-125.7 158.1 65.5 8.7 -3.6 20 21 A P B -K 100 0C 0 0, 0.0 80,-0.3 0, 0.0 79,-0.1 -0.439 38.8-121.2 -71.9 156.9 68.1 8.5 -6.4 21 22 A K S S+ 0 0 69 78,-2.5 79,-0.1 77,-0.2 78,-0.1 0.909 87.1 14.4 -69.5 -50.2 66.6 9.5 -9.8 22 23 A H S S+ 0 0 70 77,-0.3 2,-0.4 1,-0.2 -1,-0.1 0.879 114.5 35.8-100.2 -44.9 67.1 6.6 -12.2 23 24 A R > - 0 0 78 34,-0.1 3,-1.1 1,-0.1 4,-0.2 -0.923 53.4-144.5-122.9 143.8 68.0 3.2 -10.7 24 25 A R T 3 S+ 0 0 53 -2,-0.4 3,-0.1 1,-0.3 35,-0.1 0.489 105.1 55.5 -77.2 -2.2 67.1 1.5 -7.4 25 26 A N T 3 S+ 0 0 94 1,-0.1 -1,-0.3 33,-0.1 3,-0.2 -0.073 74.9 110.3-118.2 32.2 70.6 0.1 -7.3 26 27 A T < + 0 0 7 -3,-1.1 2,-1.5 31,-0.7 32,-0.2 0.946 66.4 58.3 -72.1 -50.8 72.4 3.5 -7.5 27 28 A L S S+ 0 0 2 30,-2.1 2,-0.3 29,-0.2 -1,-0.2 -0.588 79.2 124.5 -88.6 82.3 73.8 3.7 -3.9 28 29 A V >> - 0 0 49 -2,-1.5 3,-1.4 -3,-0.2 4,-1.0 -0.889 61.2 -15.2-134.7 163.5 76.0 0.7 -3.6 29 30 A N H 3> S- 0 0 116 -2,-0.3 4,-2.1 1,-0.3 3,-0.3 -0.109 130.2 -0.5 46.6-128.8 79.6 -0.2 -2.8 30 31 A E H 3> S+ 0 0 125 1,-0.2 4,-2.5 2,-0.2 -1,-0.3 0.830 134.6 56.7 -58.9 -35.3 82.0 2.8 -2.9 31 32 A I H <> S+ 0 0 20 -3,-1.4 4,-2.0 2,-0.2 -1,-0.2 0.911 108.9 44.9 -64.4 -42.6 79.2 5.1 -3.9 32 33 A A H X S+ 0 0 19 -4,-1.0 4,-2.1 -3,-0.3 -2,-0.2 0.895 112.8 52.1 -67.6 -41.2 77.1 4.2 -0.9 33 34 A E H X S+ 0 0 117 -4,-2.1 4,-1.7 1,-0.2 -2,-0.2 0.921 112.2 44.3 -61.0 -46.5 80.2 4.5 1.4 34 35 A Y H X S+ 0 0 45 -4,-2.5 4,-2.8 2,-0.2 5,-0.2 0.875 109.8 56.8 -67.3 -37.6 81.1 8.0 0.1 35 36 A T H X S+ 0 0 0 -4,-2.0 4,-2.3 1,-0.2 5,-0.2 0.922 107.1 48.0 -59.9 -44.8 77.4 9.1 0.3 36 37 A K H X S+ 0 0 75 -4,-2.1 4,-1.9 1,-0.2 -1,-0.2 0.903 112.2 51.6 -62.8 -40.0 77.3 8.3 4.0 37 38 A E H X S+ 0 0 106 -4,-1.7 4,-1.9 1,-0.2 -2,-0.2 0.935 111.4 44.3 -62.8 -48.8 80.6 10.1 4.5 38 39 A V H X S+ 0 0 5 -4,-2.8 4,-2.5 1,-0.2 -1,-0.2 0.863 111.5 52.9 -67.3 -36.5 79.5 13.3 2.7 39 40 A L H X S+ 0 0 0 -4,-2.3 4,-2.1 -5,-0.2 -1,-0.2 0.905 110.2 48.5 -66.2 -37.5 76.1 13.4 4.5 40 41 A K H X S+ 0 0 88 -4,-1.9 4,-2.1 -5,-0.2 -2,-0.2 0.883 111.6 50.6 -67.7 -37.0 77.8 13.1 7.8 41 42 A S H X S+ 0 0 80 -4,-1.9 4,-2.6 2,-0.2 -2,-0.2 0.895 109.5 49.6 -67.2 -41.1 80.2 15.9 6.9 42 43 A I H X S+ 0 0 1 -4,-2.5 4,-2.2 2,-0.2 6,-0.2 0.903 109.0 52.8 -64.9 -40.1 77.4 18.2 5.8 43 44 A A H X>S+ 0 0 3 -4,-2.1 5,-3.0 1,-0.2 4,-0.7 0.939 111.7 46.0 -59.5 -45.9 75.6 17.5 9.1 44 45 A E H ><5S+ 0 0 146 -4,-2.1 3,-0.9 1,-0.2 -2,-0.2 0.908 110.0 55.0 -62.4 -42.5 78.7 18.5 11.0 45 46 A E H 3<5S+ 0 0 85 -4,-2.6 -1,-0.2 1,-0.3 -2,-0.2 0.861 111.9 43.2 -57.9 -40.4 79.1 21.5 8.7 46 47 A L H 3<5S- 0 0 8 -4,-2.2 -1,-0.3 -5,-0.1 -2,-0.2 0.488 115.8-111.5 -86.7 -5.1 75.6 22.7 9.6 47 48 A G T <<5 + 0 0 43 -3,-0.9 2,-0.3 -4,-0.7 -3,-0.2 0.762 69.8 143.4 80.0 24.4 75.9 22.0 13.3 48 49 A C < - 0 0 13 -5,-3.0 2,-0.6 -6,-0.2 -1,-0.3 -0.684 45.6-137.7 -98.8 153.3 73.4 19.2 13.1 49 50 A E E -H 65 0B 106 16,-3.0 16,-3.1 -2,-0.3 2,-0.4 -0.944 16.3-135.8-113.0 117.4 73.5 15.9 15.0 50 51 A I E +H 64 0B 45 -2,-0.6 14,-0.3 14,-0.2 3,-0.1 -0.581 21.7 179.7 -74.4 125.3 72.6 12.8 13.1 51 52 A I E S- 0 0 78 12,-3.0 2,-0.3 1,-0.4 13,-0.2 0.834 73.9 -10.2 -89.3 -42.8 70.3 10.5 15.2 52 53 A A E -H 63 0B 33 11,-2.1 11,-3.3 2,-0.0 -1,-0.4 -0.970 61.3-172.6-157.9 141.2 70.1 7.9 12.5 53 54 A L E -H 62 0B 17 -2,-0.3 2,-0.5 9,-0.2 9,-0.2 -0.949 0.5-174.3-143.0 119.7 71.1 7.6 8.9 54 55 A E E -H 61 0B 77 7,-2.7 7,-2.3 -2,-0.4 2,-0.7 -0.963 3.7-170.6-115.2 121.7 70.4 4.8 6.4 55 56 A V E +H 60 0B 26 -2,-0.5 5,-0.2 5,-0.2 -19,-0.1 -0.901 20.8 173.3-111.7 97.4 72.0 4.9 3.0 56 57 A M E > -H 59 0B 88 3,-2.5 3,-1.1 -2,-0.7 -29,-0.2 -0.647 46.8-101.3-103.0 162.4 70.3 2.2 1.0 57 58 A P T 3 S+ 0 0 52 0, 0.0 -30,-2.1 0, 0.0 -31,-0.7 0.841 120.2 25.9 -49.6 -38.5 70.6 1.4 -2.7 58 59 A D T 3 S+ 0 0 28 -33,-0.2 2,-0.3 -32,-0.2 -33,-0.1 0.239 125.9 17.0-114.6 12.1 67.3 3.1 -3.5 59 60 A H E < - H 0 56B 29 -3,-1.1 -3,-2.5 -40,-0.2 2,-0.4 -0.977 55.1-131.6-173.0 163.1 66.8 5.7 -0.8 60 61 A I E -FH 18 55B 0 -42,-2.6 -42,-1.6 -2,-0.3 2,-0.5 -0.984 13.3-169.6-123.7 132.2 68.2 7.8 2.0 61 62 A H E -FH 17 54B 18 -7,-2.3 -7,-2.7 -2,-0.4 2,-0.5 -0.971 11.0-176.9-122.1 112.7 66.6 8.1 5.4 62 63 A L E -FH 16 53B 0 -46,-2.9 -46,-2.9 -2,-0.5 2,-0.6 -0.953 7.9-174.0-114.7 117.8 68.2 10.9 7.4 63 64 A F E +FH 15 52B 36 -11,-3.3 -12,-3.0 -2,-0.5 -11,-2.1 -0.945 23.2 167.2-112.9 111.6 67.0 11.5 11.0 64 65 A V E -FH 14 50B 0 -50,-2.5 -50,-2.5 -2,-0.6 2,-0.5 -0.945 38.8-138.6-135.8 149.9 68.7 14.6 12.4 65 66 A N E -FH 13 49B 7 -16,-3.1 -16,-3.0 -2,-0.3 -52,-0.2 -0.896 38.5-179.0-101.3 125.7 68.6 17.0 15.3 66 67 A C - 0 0 0 -54,-3.0 -54,-0.2 -2,-0.5 -18,-0.1 -0.888 32.5 -94.1-132.4 160.5 69.2 20.6 14.0 67 68 A P > - 0 0 2 0, 0.0 3,-1.4 0, 0.0 137,-0.1 -0.359 39.7-119.0 -65.9 147.6 69.5 24.2 15.1 68 69 A P T 3 S+ 0 0 3 0, 0.0 136,-2.7 0, 0.0 137,-0.4 0.567 102.7 80.7 -66.6 -7.7 66.2 26.2 14.9 69 70 A R T 3 S+ 0 0 142 134,-0.2 2,-0.4 135,-0.1 133,-0.2 0.617 80.3 82.9 -75.0 -11.1 67.8 28.7 12.4 70 71 A Y S < S- 0 0 31 -3,-1.4 134,-0.1 131,-0.1 131,-0.1 -0.801 71.9-138.5-103.8 135.0 67.2 26.2 9.5 71 72 A A > - 0 0 14 -2,-0.4 4,-2.2 131,-0.2 3,-0.5 -0.621 25.0-123.1 -82.7 142.7 64.0 25.6 7.5 72 73 A P H > S+ 0 0 0 0, 0.0 4,-2.5 0, 0.0 5,-0.2 0.895 113.0 52.1 -52.8 -41.4 63.3 22.0 6.8 73 74 A S H > S+ 0 0 19 126,-2.8 4,-2.3 1,-0.2 5,-0.2 0.842 106.6 52.6 -66.5 -32.6 63.3 22.7 3.1 74 75 A Y H > S+ 0 0 102 -3,-0.5 4,-2.1 125,-0.4 -1,-0.2 0.925 110.5 48.4 -67.4 -43.2 66.6 24.4 3.3 75 76 A L H X S+ 0 0 0 -4,-2.2 4,-2.9 2,-0.2 5,-0.3 0.945 111.6 48.6 -61.8 -48.3 68.1 21.3 5.0 76 77 A A H X S+ 0 0 0 -4,-2.5 4,-2.4 1,-0.2 5,-0.2 0.942 115.2 44.5 -58.0 -48.3 66.6 18.9 2.5 77 78 A N H X S+ 0 0 58 -4,-2.3 4,-2.4 2,-0.2 -1,-0.2 0.799 115.7 47.2 -69.4 -27.7 67.9 20.9 -0.5 78 79 A Y H X S+ 0 0 62 -4,-2.1 4,-2.6 2,-0.2 5,-0.5 0.921 113.8 45.6 -79.0 -45.7 71.3 21.5 1.0 79 80 A F H X S+ 0 0 0 -4,-2.9 4,-1.4 -5,-0.2 5,-0.2 0.932 122.2 39.8 -61.3 -44.0 71.9 17.9 2.1 80 81 A K H X S+ 0 0 11 -4,-2.4 4,-2.0 -5,-0.3 -2,-0.2 0.929 118.7 45.6 -71.2 -46.0 70.6 16.8 -1.3 81 82 A G H X S+ 0 0 43 -4,-2.4 4,-1.8 -5,-0.2 -3,-0.2 0.908 117.2 40.6 -66.4 -45.8 72.3 19.5 -3.4 82 83 A K H X S+ 0 0 46 -4,-2.6 4,-2.2 2,-0.2 5,-0.2 0.880 115.4 50.7 -73.7 -36.7 75.8 19.4 -1.8 83 84 A S H X S+ 0 0 0 -4,-1.4 4,-3.2 -5,-0.5 5,-0.3 0.920 109.9 51.9 -66.2 -39.8 75.9 15.6 -1.6 84 85 A A H X S+ 0 0 14 -4,-2.0 4,-3.0 2,-0.2 5,-0.3 0.927 109.2 49.5 -60.4 -46.2 74.9 15.3 -5.2 85 86 A R H X S+ 0 0 143 -4,-1.8 4,-1.4 1,-0.2 -1,-0.2 0.930 116.5 41.6 -58.9 -46.9 77.7 17.7 -6.3 86 87 A L H X S+ 0 0 40 -4,-2.2 4,-1.9 2,-0.2 -2,-0.2 0.901 117.0 47.3 -68.7 -42.4 80.3 15.8 -4.3 87 88 A I H X S+ 0 0 0 -4,-3.2 4,-2.3 1,-0.2 -2,-0.2 0.916 112.0 49.1 -67.0 -44.0 79.0 12.4 -5.3 88 89 A L H < S+ 0 0 54 -4,-3.0 -1,-0.2 -5,-0.3 6,-0.2 0.829 111.3 51.9 -64.7 -31.4 78.8 13.2 -9.0 89 90 A K H < S+ 0 0 169 -4,-1.4 -1,-0.2 -5,-0.3 -2,-0.2 0.885 113.7 41.8 -72.9 -38.6 82.3 14.6 -8.9 90 91 A K H < S+ 0 0 108 -4,-1.9 -2,-0.2 1,-0.3 -1,-0.2 0.782 122.2 40.7 -79.9 -26.8 83.8 11.5 -7.3 91 92 A F >< - 0 0 23 -4,-2.3 3,-2.3 -5,-0.2 -1,-0.3 -0.729 64.1-179.8-126.2 84.0 81.8 9.1 -9.5 92 93 A P G > S+ 0 0 96 0, 0.0 3,-1.3 0, 0.0 -1,-0.1 0.677 76.8 71.2 -53.6 -23.3 81.7 10.4 -13.1 93 94 A Q G 3 S+ 0 0 120 1,-0.3 3,-0.2 2,-0.1 -5,-0.1 0.666 79.2 73.9 -71.7 -14.7 79.6 7.4 -14.2 94 95 A L G < + 0 0 11 -3,-2.3 -1,-0.3 -6,-0.2 -6,-0.1 0.392 67.7 125.6 -77.5 7.1 76.6 8.8 -12.3 95 96 A N < + 0 0 122 -3,-1.3 -1,-0.2 1,-0.2 -2,-0.1 0.763 24.0 107.2 -19.2 -99.2 76.7 11.1 -15.2 96 97 A K S S- 0 0 127 -3,-0.2 2,-0.2 3,-0.0 -1,-0.2 0.492 95.3 -86.6 19.3 34.5 73.0 10.7 -16.4 97 98 A G S S+ 0 0 45 -76,-0.0 -76,-0.0 2,-0.0 2,-0.0 -0.490 94.9 17.9 84.6-145.2 71.9 14.1 -15.2 98 99 A K - 0 0 150 -2,-0.2 -77,-0.2 1,-0.1 4,-0.0 -0.311 59.4-145.3 -64.8 146.0 70.7 14.8 -11.6 99 100 A L S S+ 0 0 11 -78,-0.1 -78,-2.5 -19,-0.1 -77,-0.3 0.890 82.0 35.7 -78.8 -42.1 71.6 12.3 -8.9 100 101 A W B S-K 20 0C 16 -80,-0.3 2,-0.1 -79,-0.1 -2,-0.0 -0.682 85.2-113.0-109.3 164.1 68.4 12.8 -6.9 101 102 A T - 0 0 8 -82,-1.6 -1,-0.1 -2,-0.2 -80,-0.1 -0.369 37.3-109.3 -82.0 174.5 64.8 13.4 -8.0 102 103 A R S S+ 0 0 135 -2,-0.1 2,-0.1 -81,-0.1 36,-0.1 0.559 90.1 87.9 -85.4 -5.7 63.4 16.8 -7.0 103 104 A S + 0 0 2 -84,-0.1 -84,-0.3 34,-0.1 2,-0.3 -0.432 48.1 167.8 -92.8 165.5 61.1 15.4 -4.3 104 105 A Y E - 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