==== Secondary Structure Definition by the program DSSP, updated CMBI version by ElmK / April 1,2000 ==== DATE=20-JUL-2011 . REFERENCE W. KABSCH AND C.SANDER, BIOPOLYMERS 22 (1983) 2577-2637 . HEADER OXIDOREDUCTASE 06-DEC-05 2F9Q . COMPND 2 MOLECULE: CYTOCHROME P450 2D6; . SOURCE 2 ORGANISM_SCIENTIFIC: HOMO SAPIENS; . AUTHOR P.ROWLAND . 1810 8 0 0 0 TOTAL NUMBER OF RESIDUES, NUMBER OF CHAINS, NUMBER OF SS-BRIDGES(TOTAL,INTRACHAIN,INTERCHAIN) . 71265.0 ACCESSIBLE SURFACE OF PROTEIN (ANGSTROM**2) . 1226 67.7 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(J) , SAME NUMBER PER 100 RESIDUES . 20 1.1 TOTAL NUMBER OF HYDROGEN BONDS IN PARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 101 5.6 TOTAL NUMBER OF HYDROGEN BONDS IN ANTIPARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-5), SAME NUMBER PER 100 RESIDUES . 4 0.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-4), SAME NUMBER PER 100 RESIDUES . 4 0.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-3), SAME NUMBER PER 100 RESIDUES . 4 0.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-2), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-1), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+0), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+1), SAME NUMBER PER 100 RESIDUES . 69 3.8 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+2), SAME NUMBER PER 100 RESIDUES . 201 11.1 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+3), SAME NUMBER PER 100 RESIDUES . 747 41.3 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+4), SAME NUMBER PER 100 RESIDUES . 33 1.8 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+5), SAME NUMBER PER 100 RESIDUES . 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 *** HISTOGRAMS OF *** . 0 0 0 0 8 1 0 4 0 0 6 2 1 8 0 0 4 11 1 3 0 3 1 0 0 0 0 0 0 4 RESIDUES PER ALPHA HELIX . 0 0 0 4 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PARALLEL BRIDGES PER LADDER . 12 7 9 4 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 ANTIPARALLEL BRIDGES PER LADDER . 12 0 0 4 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 LADDERS PER SHEET . # RESIDUE AA STRUCTURE BP1 BP2 ACC N-H-->O O-->H-N N-H-->O O-->H-N TCO KAPPA ALPHA PHI PSI X-CA Y-CA Z-CA 1 34 A P 0 0 45 0, 0.0 26,-0.0 0, 0.0 343,-0.0 0.000 360.0 360.0 360.0 72.7 -8.9 52.8 -35.2 2 35 A P + 0 0 41 0, 0.0 25,-0.1 0, 0.0 22,-0.1 0.590 360.0 127.1 -21.1-143.4 -11.5 50.1 -36.6 3 36 A G - 0 0 5 16,-0.1 26,-0.1 23,-0.1 2,-0.1 0.190 56.7 -65.3 97.3 152.1 -15.3 50.0 -35.7 4 37 A P - 0 0 44 0, 0.0 27,-0.5 0, 0.0 26,-0.2 -0.332 36.2-144.5 -84.3 152.5 -18.7 49.9 -37.6 5 38 A L - 0 0 65 1,-0.1 -2,-0.0 26,-0.1 24,-0.0 -0.754 23.2-121.0-114.4 154.5 -20.3 52.5 -39.9 6 39 A P + 0 0 45 0, 0.0 -1,-0.1 0, 0.0 1533,-0.1 0.063 50.2 148.6 -96.5-173.9 -24.0 53.3 -40.1 7 40 A L 0 0 84 1531,-0.1 1530,-0.1 1530,-0.1 1531,-0.0 -0.007 360.0 360.0 178.3 22.4 -27.1 53.5 -42.4 8 41 A P 0 0 96 0, 0.0 1530,-0.0 0, 0.0 1531,-0.0 0.345 360.0 360.0 -77.4 360.0 -29.1 52.5 -39.4 9 ! 0 0 0 0, 0.0 0, 0.0 0, 0.0 0, 0.0 0.000 360.0 360.0 360.0 360.0 0.0 0.0 0.0 10 52 A Q 0 0 230 0, 0.0 2,-0.3 0, 0.0 0, 0.0 0.000 360.0 360.0 360.0 -25.4 -30.1 48.5 -23.8 11 53 A N - 0 0 68 1,-0.1 429,-0.1 2,-0.0 0, 0.0 -0.878 360.0-175.3-112.4 149.0 -27.5 46.0 -22.5 12 54 A T > + 0 0 37 -2,-0.3 4,-2.1 3,-0.1 5,-0.1 0.499 53.4 72.4-132.5 -13.4 -24.0 45.9 -23.9 13 55 A P H > S+ 0 0 18 0, 0.0 4,-1.2 0, 0.0 5,-0.1 0.943 101.4 46.7 -89.2 -46.1 -21.3 43.3 -23.0 14 56 A Y H > S+ 0 0 76 1,-0.2 4,-0.7 2,-0.2 -2,-0.1 0.824 115.9 55.9 -55.6 -28.1 -22.5 40.1 -24.8 15 57 A C H >> S+ 0 0 59 1,-0.2 3,-1.5 2,-0.2 4,-1.2 0.978 102.6 48.7 -64.7 -59.8 -22.8 42.8 -27.5 16 58 A F H 3X S+ 0 0 9 -4,-2.1 4,-2.0 1,-0.3 -1,-0.2 0.658 99.8 69.8 -54.7 -20.2 -19.1 44.1 -27.5 17 59 A D H 3X S+ 0 0 29 -4,-1.2 4,-2.2 2,-0.2 -1,-0.3 0.903 99.5 47.4 -68.5 -36.7 -18.1 40.4 -27.7 18 60 A Q H << S+ 0 0 10 -3,-1.5 -2,-0.2 -4,-0.7 5,-0.2 0.945 110.6 49.5 -64.4 -52.1 -19.4 40.3 -31.3 19 61 A L H >X S+ 0 0 31 -4,-1.2 4,-2.4 1,-0.2 3,-0.9 0.855 110.4 53.5 -60.1 -35.5 -17.7 43.6 -32.3 20 62 A R H 3X S+ 0 0 43 -4,-2.0 4,-2.7 1,-0.2 -1,-0.2 0.954 110.2 45.0 -58.7 -52.5 -14.5 42.2 -30.8 21 63 A R H 3< S+ 0 0 64 -4,-2.2 -1,-0.2 1,-0.2 -2,-0.2 0.228 120.6 42.2 -83.0 14.2 -14.8 39.0 -32.9 22 64 A R H <4 S+ 0 0 91 -3,-0.9 -1,-0.2 2,-0.2 -2,-0.2 0.534 131.6 19.3-119.6 -26.5 -15.6 41.1 -36.0 23 65 A F H < S- 0 0 52 -4,-2.4 -3,-0.2 1,-0.4 -2,-0.2 0.418 104.9-112.6-125.5 -5.5 -13.1 43.9 -35.5 24 66 A G < - 0 0 36 -4,-2.7 -1,-0.4 -5,-0.2 -2,-0.2 -0.406 53.2 -49.6 86.6-171.5 -10.4 42.7 -33.2 25 67 A D S S+ 0 0 45 -2,-0.1 2,-0.5 1,-0.1 15,-0.2 0.896 122.6 49.3 -75.4 -49.0 -9.6 43.9 -29.7 26 68 A V S S+ 0 0 4 13,-0.2 2,-0.3 -3,-0.2 13,-0.2 -0.844 74.9 157.0 -89.9 127.3 -9.5 47.6 -30.0 27 69 A F E -A 38 0A 1 11,-1.5 11,-2.4 -2,-0.5 2,-0.4 -0.960 33.0-115.3-146.5 167.6 -12.4 49.1 -31.9 28 70 A S E -A 37 0A 0 -2,-0.3 2,-0.4 9,-0.2 3,-0.3 -0.878 10.7-160.0-113.1 135.1 -14.4 52.3 -32.3 29 71 A L E S+A 36 0A 36 7,-4.2 7,-2.9 -2,-0.4 -24,-0.1 -0.835 82.7 10.8-101.1 145.9 -17.9 53.4 -31.5 30 72 A Q S > S+ 0 0 61 -2,-0.4 3,-1.5 5,-0.2 -1,-0.2 0.793 72.1 173.0 61.0 36.0 -19.4 56.4 -33.3 31 73 A L T 3 S+ 0 0 20 -27,-0.5 1519,-0.3 -3,-0.3 -26,-0.1 0.821 74.5 48.4 -41.5 -44.5 -16.6 56.6 -35.8 32 74 A A T 3 S+ 0 0 5 2,-0.1 -1,-0.3 1521,-0.1 1521,-0.1 0.762 120.7 31.9 -75.4 -25.6 -18.3 59.3 -37.8 33 75 A W S < S- 0 0 13 -3,-1.5 1517,-1.3 1,-0.2 -2,-0.1 0.769 134.8 -31.3 -94.0-103.6 -19.3 61.6 -34.9 34 76 A T S S- 0 0 45 1515,-0.2 -1,-0.2 1514,-0.1 -2,-0.1 -0.998 75.4-120.8-112.6 126.3 -16.8 61.5 -32.0 35 77 A P - 0 0 22 0, 0.0 317,-0.6 0, 0.0 2,-0.4 -0.365 21.6-163.4 -63.2 137.3 -15.1 58.1 -31.7 36 78 A V E -Ab 29 352A 2 -7,-2.9 -7,-4.2 315,-0.1 2,-0.5 -0.998 16.7-145.2-118.8 126.0 -15.5 56.2 -28.5 37 79 A V E -Ab 28 353A 3 315,-2.9 317,-3.0 -2,-0.4 2,-0.6 -0.827 6.8-155.4-105.1 131.2 -12.9 53.4 -28.2 38 80 A V E -Ab 27 354A 0 -11,-2.4 -11,-1.5 -2,-0.5 2,-0.4 -0.901 9.3-151.8-103.5 120.4 -13.5 50.1 -26.5 39 81 A L E + b 0 355A 0 315,-3.3 317,-2.0 -2,-0.6 -13,-0.2 -0.773 22.7 165.9 -94.5 133.0 -10.4 48.2 -25.2 40 82 A N + 0 0 0 -2,-0.4 -1,-0.1 -15,-0.2 -14,-0.1 0.826 39.9 72.3-121.1 -45.2 -10.8 44.5 -25.2 41 83 A G S > S- 0 0 3 1,-0.2 4,-1.2 319,-0.0 319,-0.1 -0.062 74.1-125.5 -56.5 169.5 -7.6 42.3 -24.7 42 84 A L H > S+ 0 0 25 2,-0.2 4,-3.0 317,-0.1 5,-0.3 0.796 125.6 53.8 -64.7 -41.5 -5.8 42.0 -21.4 43 85 A A H > S+ 0 0 44 1,-0.2 4,-2.0 2,-0.2 5,-0.1 0.899 108.4 45.1 -52.8 -48.5 -3.3 43.0 -24.0 44 86 A A H > S+ 0 0 0 2,-0.2 4,-1.4 1,-0.2 -1,-0.2 0.914 114.8 47.3 -65.7 -44.4 -5.3 46.0 -25.1 45 87 A V H X S+ 0 0 0 -4,-1.2 4,-2.7 2,-0.2 5,-0.5 0.911 111.8 49.7 -64.3 -45.0 -6.1 47.2 -21.6 46 88 A R H X>S+ 0 0 100 -4,-3.0 4,-2.1 1,-0.2 5,-1.6 0.943 109.4 52.7 -61.8 -45.6 -2.6 46.9 -20.3 47 89 A E H <>S+ 0 0 42 -4,-2.0 5,-1.4 -5,-0.3 -1,-0.2 0.829 116.3 37.7 -60.8 -34.1 -1.3 48.8 -23.2 48 90 A A H <5S+ 0 0 1 -4,-1.4 7,-0.2 3,-0.2 -2,-0.2 0.901 125.8 31.7 -84.4 -42.3 -3.6 51.8 -22.7 49 91 A L H <5S+ 0 0 2 -4,-2.7 349,-0.9 -5,-0.2 350,-0.2 0.786 132.4 25.7 -96.4 -30.5 -3.8 52.0 -18.9 50 92 A V T ><5S+ 0 0 76 -4,-2.1 3,-1.7 -5,-0.5 4,-0.3 0.861 126.2 39.5 -93.1 -69.2 -0.4 50.9 -17.9 51 93 A T T 3 >> - 0 0 6 17,-0.1 4,-0.9 -3,-0.1 3,-0.5 -0.988 36.9-122.2-144.2 146.2 -20.2 64.4 -16.9 63 105 A P H 3> S+ 0 0 0 0, 0.0 4,-3.2 0, 0.0 5,-0.3 0.753 103.8 78.0 -60.7 -18.5 -23.1 66.6 -15.9 64 106 A I H 3> S+ 0 0 0 2,-0.2 4,-3.0 1,-0.2 5,-0.2 0.955 92.8 47.9 -51.6 -57.4 -22.7 65.2 -12.3 65 107 A T H <4>S+ 0 0 37 -3,-0.5 5,-2.9 1,-0.2 -1,-0.2 0.907 117.2 42.5 -46.5 -50.9 -19.6 67.4 -11.6 66 108 A Q H ><5S+ 0 0 64 -4,-0.9 3,-1.3 3,-0.2 -1,-0.2 0.895 113.8 50.3 -69.0 -41.4 -21.3 70.5 -12.9 67 109 A I H 3<5S+ 0 0 23 -4,-3.2 -2,-0.2 1,-0.3 -1,-0.2 0.938 110.4 49.4 -65.0 -45.8 -24.7 69.8 -11.2 68 110 A L T 3<5S- 0 0 18 -4,-3.0 -1,-0.3 -5,-0.3 -2,-0.2 0.324 117.0-118.7 -74.1 8.5 -23.1 69.1 -7.8 69 111 A G T < 5 + 0 0 52 -3,-1.3 -3,-0.2 -5,-0.2 -2,-0.1 0.920 55.8 160.3 54.2 53.9 -21.3 72.4 -8.4 70 112 A F < + 0 0 14 -5,-2.9 3,-0.1 -6,-0.1 -4,-0.1 0.940 13.9 132.4 -70.0 -93.3 -17.8 70.9 -8.3 71 113 A G > + 0 0 45 1,-0.2 3,-1.9 2,-0.1 -1,-0.1 -0.402 49.9 24.8 76.3-146.1 -15.3 73.2 -9.9 72 114 A P T 3 S- 0 0 87 0, 0.0 -1,-0.2 0, 0.0 2,-0.2 0.618 130.6 -25.8 -44.0 -90.2 -12.0 74.5 -9.0 73 115 A R T 3 S+ 0 0 89 173,-0.1 176,-0.3 1,-0.1 179,-0.2 -0.540 114.8 99.5-112.4 57.8 -10.2 72.2 -6.6 74 116 A S < + 0 0 3 -3,-1.9 2,-0.3 -2,-0.2 -1,-0.1 -0.293 48.3 107.6-142.8 47.8 -13.4 70.7 -5.2 75 117 A Q - 0 0 31 1,-0.3 181,-0.1 -5,-0.1 180,-0.1 -0.828 62.3-130.0-117.5 162.8 -13.8 67.5 -7.1 76 118 A G - 0 0 0 -2,-0.3 -1,-0.3 179,-0.1 180,-0.2 0.627 33.6 -93.8 -84.3-119.5 -13.3 64.1 -5.4 77 119 A V S S+ 0 0 12 178,-0.2 3,-0.4 179,-0.1 13,-0.1 0.555 111.7 60.8-121.5 -49.7 -11.1 61.4 -6.8 78 120 A F S S+ 0 0 67 1,-0.3 178,-0.0 178,-0.1 8,-0.0 0.835 123.7 15.7 -51.4 -52.4 -13.6 59.3 -8.7 79 121 A L S S+ 0 0 13 2,-0.1 -1,-0.3 -20,-0.1 -20,-0.1 0.351 96.0 126.0-106.8 1.2 -14.9 61.9 -11.3 80 122 A A - 0 0 0 -3,-0.4 2,-0.3 1,-0.1 -21,-0.1 -0.277 63.9-115.8 -60.4 146.5 -12.0 64.4 -10.7 81 123 A R > - 0 0 89 -23,-0.1 5,-0.8 1,-0.1 -23,-0.1 -0.654 36.2 -90.8 -88.4 143.6 -10.3 65.3 -13.9 82 124 A Y T 5S+ 0 0 49 -2,-0.3 -1,-0.1 4,-0.1 -24,-0.1 -0.359 86.5 92.6 -51.0 119.6 -6.7 64.5 -14.4 83 125 A G T >5S- 0 0 13 1,-0.1 4,-2.3 -2,-0.1 5,-0.2 -0.936 95.7 -55.0 175.9-175.1 -4.7 67.5 -13.0 84 126 A P H >5S+ 0 0 88 0, 0.0 4,-3.2 0, 0.0 5,-0.2 0.914 128.0 58.5 -42.8 -54.4 -3.1 68.8 -9.8 85 127 A A H >5S+ 0 0 14 1,-0.2 4,-1.2 2,-0.2 162,-0.1 0.874 111.0 37.0 -45.1 -61.8 -6.4 68.2 -7.9 86 128 A W H >X S+ 0 0 160 -4,-2.1 4,-1.6 -5,-0.2 3,-0.6 0.992 111.2 47.1 -66.9 -64.5 -1.2 52.4 3.7 99 141 A N H 3< S+ 0 0 82 -4,-2.3 5,-0.3 1,-0.3 7,-0.2 0.814 120.6 37.4 -48.2 -44.9 -0.4 54.0 7.1 100 142 A L H 3< S+ 0 0 19 -4,-1.8 7,-2.3 -5,-0.2 8,-0.4 0.782 111.9 61.6 -78.3 -29.5 -3.7 52.9 8.7 101 143 A G H << S+ 0 0 10 -4,-1.2 7,-0.7 -3,-0.6 -2,-0.2 0.985 130.1 1.9 -57.2 -67.2 -3.7 49.5 6.8 102 144 A L S >< S+ 0 0 111 -4,-1.6 3,-2.7 3,-0.1 4,-0.3 0.930 109.9 89.7 -75.3 -89.6 -0.5 48.2 8.4 103 145 A G T 3 S- 0 0 57 1,-0.3 -3,-0.2 -5,-0.2 -4,-0.1 0.099 115.3 -13.1 28.0 -84.0 0.7 51.0 10.9 104 146 A K T 3 S- 0 0 192 -5,-0.3 -1,-0.3 -4,-0.3 -2,-0.1 -0.287 99.8 -92.7-136.2 51.6 -1.0 50.0 14.2 105 147 A K S <> S+ 0 0 148 -3,-2.7 4,-3.5 1,-0.1 5,-0.2 0.671 82.2 143.4 47.6 22.3 -3.5 47.4 12.9 106 148 A S H > S+ 0 0 26 -4,-0.3 4,-2.6 -6,-0.3 5,-0.2 0.974 71.6 35.6 -53.7 -67.8 -6.0 50.4 12.6 107 149 A L H > S+ 0 0 2 -7,-2.3 4,-2.6 1,-0.2 5,-0.2 0.914 119.5 51.2 -55.6 -47.1 -7.8 49.3 9.5 108 150 A E H > S+ 0 0 40 -7,-0.7 4,-3.3 -8,-0.4 5,-0.3 0.932 109.8 50.5 -52.9 -47.5 -7.6 45.6 10.5 109 151 A Q H X S+ 0 0 109 -4,-3.5 4,-2.3 1,-0.2 5,-0.3 0.961 111.0 47.9 -59.4 -53.8 -9.0 46.4 13.9 110 152 A W H X S+ 0 0 15 -4,-2.6 4,-1.4 -5,-0.2 -1,-0.2 0.881 116.2 44.2 -54.8 -44.2 -11.9 48.3 12.5 111 153 A V H X S+ 0 0 0 -4,-2.6 4,-2.7 -5,-0.2 5,-0.2 0.971 110.8 52.2 -68.2 -53.0 -12.7 45.5 10.0 112 154 A T H X S+ 0 0 20 -4,-3.3 4,-1.7 1,-0.2 -2,-0.2 0.911 112.6 45.1 -49.6 -48.4 -12.3 42.5 12.4 113 155 A E H X S+ 0 0 76 -4,-2.3 4,-1.9 -5,-0.3 -1,-0.2 0.879 110.3 54.6 -67.5 -35.5 -14.7 44.0 14.9 114 156 A E H X S+ 0 0 2 -4,-1.4 4,-3.0 -5,-0.3 -1,-0.2 0.918 107.1 51.4 -60.8 -41.2 -17.1 44.9 12.1 115 157 A A H X S+ 0 0 1 -4,-2.7 4,-3.1 1,-0.2 -2,-0.2 0.876 105.9 56.0 -56.6 -38.5 -17.0 41.2 11.1 116 158 A A H X S+ 0 0 35 -4,-1.7 4,-1.7 -5,-0.2 -1,-0.2 0.879 110.9 43.1 -68.3 -37.8 -17.8 40.3 14.6 117 159 A C H X S+ 0 0 36 -4,-1.9 4,-3.7 2,-0.2 -2,-0.2 0.940 111.2 53.5 -71.2 -47.3 -20.9 42.4 14.5 118 160 A L H X S+ 0 0 0 -4,-3.0 4,-2.6 1,-0.2 -2,-0.2 0.925 110.4 49.5 -49.4 -46.5 -21.9 41.2 11.1 119 161 A C H X S+ 0 0 7 -4,-3.1 4,-2.4 2,-0.2 -1,-0.2 0.889 110.9 49.0 -61.5 -40.9 -21.6 37.7 12.5 120 162 A A H X S+ 0 0 49 -4,-1.7 4,-1.9 2,-0.2 -2,-0.2 0.941 110.3 51.2 -61.9 -47.8 -23.7 38.7 15.5 121 163 A A H < S+ 0 0 14 -4,-3.7 -2,-0.2 1,-0.2 4,-0.2 0.909 111.8 46.9 -55.9 -45.6 -26.3 40.2 13.1 122 164 A F H >< S+ 0 0 1 -4,-2.6 3,-2.2 1,-0.2 5,-0.3 0.940 109.3 53.7 -58.0 -49.3 -26.4 36.9 11.1 123 165 A A H >< S+ 0 0 48 -4,-2.4 3,-1.5 1,-0.3 -1,-0.2 0.788 96.5 68.4 -62.7 -30.6 -26.7 34.9 14.2 124 166 A N T 3< S+ 0 0 126 -4,-1.9 -1,-0.3 1,-0.3 -2,-0.2 0.532 86.3 68.2 -65.8 -8.8 -29.7 37.0 15.3 125 167 A H T X S- 0 0 59 -3,-2.2 3,-1.9 -4,-0.2 -1,-0.3 0.421 96.0-140.3 -90.1 0.8 -31.8 35.5 12.4 126 168 A S T < - 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