==== Secondary Structure Definition by the program DSSP, updated CMBI version by ElmK / April 1,2000 ==== DATE=16-AUG-2013 . REFERENCE W. KABSCH AND C.SANDER, BIOPOLYMERS 22 (1983) 2577-2637 . HEADER HYDROLASE 29-JUN-12 4FVL . COMPND 2 MOLECULE: COLLAGENASE 3; . SOURCE 2 ORGANISM_SCIENTIFIC: HOMO SAPIENS; . AUTHOR E.A.STURA,L.VERA,R.VISSE,H.NAGASE,V.DIVE . 770 4 1 1 0 TOTAL NUMBER OF RESIDUES, NUMBER OF CHAINS, NUMBER OF SS-BRIDGES(TOTAL,INTRACHAIN,INTERCHAIN) . 36491.0 ACCESSIBLE SURFACE OF PROTEIN (ANGSTROM**2) . 467 60.6 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(J) , SAME NUMBER PER 100 RESIDUES . 38 4.9 TOTAL NUMBER OF HYDROGEN BONDS IN PARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 177 23.0 TOTAL NUMBER OF HYDROGEN BONDS IN ANTIPARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 8 1.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-5), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-4), SAME NUMBER PER 100 RESIDUES . 14 1.8 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-3), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-2), SAME NUMBER PER 100 RESIDUES . 2 0.3 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-1), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+0), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+1), SAME NUMBER PER 100 RESIDUES . 62 8.1 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+2), SAME NUMBER PER 100 RESIDUES . 88 11.4 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+3), SAME NUMBER PER 100 RESIDUES . 87 11.3 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+4), SAME NUMBER PER 100 RESIDUES . 12 1.6 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+5), SAME NUMBER PER 100 RESIDUES . 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 *** HISTOGRAMS OF *** . 0 0 0 3 0 0 0 0 0 1 2 0 2 0 0 2 0 0 0 0 0 0 0 0 0 0 0 0 0 0 RESIDUES PER ALPHA HELIX . 2 2 2 4 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PARALLEL BRIDGES PER LADDER . 0 10 2 3 13 6 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 ANTIPARALLEL BRIDGES PER LADDER . 4 0 4 4 0 2 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 LADDERS PER SHEET . # RESIDUE AA STRUCTURE BP1 BP2 ACC N-H-->O O-->H-N N-H-->O O-->H-N TCO KAPPA ALPHA PHI PSI X-CA Y-CA Z-CA 1 104 A Y 0 0 101 0, 0.0 2,-0.4 0, 0.0 129,-0.1 0.000 360.0 360.0 360.0 164.5 -6.2 15.0 15.5 2 105 A N E -A 128 0A 76 126,-0.6 126,-4.1 135,-0.0 2,-0.2 -0.991 360.0-158.9-140.2 132.3 -9.1 13.3 17.2 3 106 A V E -A 127 0A 43 -2,-0.4 124,-0.2 124,-0.3 3,-0.1 -0.500 38.0 -90.7 -92.3 169.9 -11.8 10.9 16.1 4 107 A F > - 0 0 2 122,-1.0 3,-2.2 4,-0.2 -1,-0.1 -0.533 49.4 -88.4 -81.5 149.3 -15.2 10.4 17.8 5 108 A P G > S+ 0 0 62 0, 0.0 3,-0.6 0, 0.0 -1,-0.1 -0.280 121.0 23.3 -51.0 140.6 -15.8 7.9 20.6 6 109 A R G 3 S- 0 0 214 1,-0.2 3,-0.2 -3,-0.1 -2,-0.1 0.390 121.7-100.2 72.1 -0.7 -16.7 4.5 19.2 7 110 A T G < - 0 0 68 -3,-2.2 2,-0.7 1,-0.2 -1,-0.2 0.951 42.6-149.4 52.7 58.6 -15.0 5.8 16.1 8 111 A L < + 0 0 21 -3,-0.6 2,-0.3 -4,-0.1 -4,-0.2 -0.453 31.4 160.3 -58.6 103.4 -18.1 6.8 14.2 9 112 A K - 0 0 109 -2,-0.7 117,-0.1 -3,-0.2 116,-0.1 -0.924 44.5 -99.8-123.1 153.8 -17.1 6.3 10.5 10 113 A W - 0 0 19 115,-0.4 5,-0.0 -2,-0.3 115,-0.0 -0.346 23.9-143.5 -64.9 155.6 -19.4 5.9 7.5 11 114 A S S S+ 0 0 121 -2,-0.0 2,-0.2 2,-0.0 -1,-0.1 0.892 78.8 48.2 -87.7 -48.6 -19.9 2.4 6.2 12 115 A K - 0 0 87 2,-0.1 -2,-0.1 1,-0.1 36,-0.0 -0.494 69.3-140.6 -91.0 162.5 -20.0 3.0 2.5 13 116 A M S S+ 0 0 73 -2,-0.2 36,-2.5 1,-0.1 2,-0.8 0.761 85.1 76.9 -90.2 -30.2 -17.6 5.0 0.3 14 117 A N E +b 49 0B 111 34,-0.2 2,-0.3 36,-0.1 36,-0.2 -0.747 68.4 179.9 -88.7 110.6 -20.2 6.5 -2.0 15 118 A L E -b 50 0B 0 34,-2.9 36,-2.1 -2,-0.8 2,-0.3 -0.806 16.3-146.6-118.4 154.1 -22.0 9.3 -0.1 16 119 A T E -b 51 0B 31 -2,-0.3 44,-2.8 34,-0.2 43,-0.3 -0.886 11.9-177.7-119.2 147.4 -24.7 11.8 -0.8 17 120 A Y E -bc 52 60B 15 34,-2.5 36,-2.9 -2,-0.3 2,-0.3 -0.925 7.4-160.9-133.9 164.8 -25.5 15.4 0.2 18 121 A R E - c 0 61B 59 42,-2.1 44,-2.9 -2,-0.3 2,-1.4 -0.953 20.1-138.4-149.1 125.6 -28.3 17.9 -0.4 19 122 A I E - c 0 62B 9 34,-0.4 44,-0.2 -2,-0.3 42,-0.1 -0.685 24.2-173.1 -81.8 91.1 -28.3 21.7 -0.1 20 123 A V - 0 0 62 42,-2.0 2,-0.3 -2,-1.4 -1,-0.2 0.852 64.7 -9.6 -59.4 -40.6 -31.8 21.9 1.5 21 124 A N S S- 0 0 61 41,-0.8 2,-0.3 -3,-0.1 -1,-0.1 -0.898 70.3-126.9-147.5-179.6 -31.8 25.7 1.4 22 125 A Y - 0 0 45 -2,-0.3 6,-0.1 41,-0.1 42,-0.1 -0.889 16.1-114.1-133.4 159.1 -29.5 28.6 0.7 23 126 A T > - 0 0 3 -2,-0.3 3,-0.6 4,-0.1 8,-0.1 -0.706 16.9-140.9 -86.1 145.0 -28.2 31.9 2.1 24 127 A P T 3 S+ 0 0 114 0, 0.0 -1,-0.1 0, 0.0 4,-0.0 0.596 94.3 80.7 -73.1 -18.3 -29.2 35.2 0.3 25 128 A D T 3 S+ 0 0 68 77,-0.1 2,-0.3 75,-0.1 78,-0.0 0.758 102.9 26.7 -61.1 -29.2 -25.6 36.4 1.0 26 129 A M S < S- 0 0 12 -3,-0.6 79,-0.1 79,-0.1 2,-0.1 -0.911 90.0-102.1-130.0 160.2 -24.4 34.4 -2.0 27 130 A T > - 0 0 68 -2,-0.3 4,-1.9 1,-0.1 5,-0.2 -0.346 38.2-108.4 -70.4 162.7 -25.9 33.2 -5.3 28 131 A H H > S+ 0 0 106 2,-0.2 4,-2.4 1,-0.2 5,-0.2 0.883 118.8 54.1 -57.9 -42.7 -26.9 29.5 -5.6 29 132 A S H > S+ 0 0 80 2,-0.2 4,-2.4 1,-0.2 -1,-0.2 0.954 108.2 47.5 -59.9 -52.8 -24.0 28.8 -8.0 30 133 A E H > S+ 0 0 80 2,-0.2 4,-2.8 1,-0.2 -1,-0.2 0.895 112.7 49.9 -52.2 -47.6 -21.3 30.2 -5.5 31 134 A V H X S+ 0 0 0 -4,-1.9 4,-2.7 2,-0.2 -1,-0.2 0.925 112.1 47.4 -58.8 -46.8 -22.8 28.2 -2.6 32 135 A E H X S+ 0 0 48 -4,-2.4 4,-2.4 1,-0.2 -2,-0.2 0.862 112.2 50.8 -62.6 -36.9 -22.8 25.0 -4.7 33 136 A K H X S+ 0 0 152 -4,-2.4 4,-2.7 -5,-0.2 -2,-0.2 0.927 110.5 48.8 -65.9 -45.4 -19.2 25.8 -5.7 34 137 A A H X S+ 0 0 7 -4,-2.8 4,-2.8 2,-0.2 -2,-0.2 0.954 113.6 45.4 -60.8 -51.4 -18.2 26.3 -2.1 35 138 A F H X S+ 0 0 0 -4,-2.7 4,-2.1 1,-0.2 -1,-0.2 0.908 112.9 50.7 -60.4 -43.6 -19.8 23.0 -0.9 36 139 A K H X S+ 0 0 148 -4,-2.4 4,-1.9 -5,-0.2 -1,-0.2 0.916 112.5 47.6 -57.3 -44.2 -18.4 21.1 -3.9 37 140 A K H X S+ 0 0 76 -4,-2.7 4,-1.7 1,-0.2 -2,-0.2 0.933 108.6 54.4 -62.0 -46.1 -14.9 22.5 -3.1 38 141 A A H X S+ 0 0 0 -4,-2.8 4,-0.7 1,-0.2 -1,-0.2 0.869 110.6 45.4 -59.0 -40.3 -15.3 21.6 0.6 39 142 A F H >X S+ 0 0 4 -4,-2.1 4,-2.7 1,-0.2 3,-0.7 0.868 106.5 60.1 -70.0 -35.8 -16.1 18.0 -0.3 40 143 A K H 3X S+ 0 0 105 -4,-1.9 4,-2.2 1,-0.2 -2,-0.2 0.845 94.5 64.6 -63.5 -32.9 -13.2 17.9 -2.7 41 144 A V H 3< S+ 0 0 16 -4,-1.7 -1,-0.2 1,-0.2 -2,-0.2 0.905 113.1 32.9 -51.7 -45.6 -10.8 18.7 0.1 42 145 A W H X< S+ 0 0 0 -4,-0.7 3,-1.4 -3,-0.7 4,-0.4 0.879 116.1 54.1 -83.9 -43.5 -11.6 15.4 1.8 43 146 A S H >< S+ 0 0 33 -4,-2.7 3,-0.7 1,-0.2 -2,-0.2 0.786 98.2 67.7 -61.5 -26.4 -12.3 13.3 -1.3 44 147 A D T 3< S+ 0 0 38 -4,-2.2 -1,-0.2 -5,-0.2 122,-0.2 0.671 111.1 31.6 -68.2 -16.2 -8.8 14.2 -2.7 45 148 A V T < S+ 0 0 2 -3,-1.4 121,-0.3 -4,-0.2 -1,-0.2 0.283 119.8 52.7-126.1 8.4 -7.1 12.3 0.1 46 149 A T S < S- 0 0 6 -3,-0.7 121,-0.3 -4,-0.4 120,-0.2 -0.885 82.4-113.5-136.7 166.6 -9.6 9.4 0.6 47 150 A P S S+ 0 0 49 0, 0.0 -4,-0.1 0, 0.0 119,-0.1 0.551 79.4 123.3 -75.2 -7.9 -11.6 6.7 -1.3 48 151 A L - 0 0 2 -6,-0.2 2,-0.4 -34,-0.1 -2,-0.2 -0.179 46.1-161.6 -57.9 144.6 -14.7 8.7 -0.4 49 152 A N E -b 14 0B 93 -36,-2.5 -34,-2.9 2,-0.0 2,-0.4 -0.985 2.8-151.2-129.9 140.1 -17.0 10.0 -3.2 50 153 A F E -b 15 0B 34 -2,-0.4 2,-0.4 -36,-0.2 -34,-0.2 -0.937 10.6-176.4-121.6 135.1 -19.6 12.7 -3.0 51 154 A T E -b 16 0B 78 -36,-2.1 -34,-2.5 -2,-0.4 2,-0.4 -0.987 19.3-136.2-130.7 133.0 -22.9 13.1 -5.0 52 155 A R E -b 17 0B 64 -2,-0.4 2,-0.4 -36,-0.2 -34,-0.2 -0.709 15.2-160.2 -91.4 132.7 -25.3 16.0 -4.8 53 156 A L - 0 0 50 -36,-2.9 -34,-0.4 -2,-0.4 5,-0.1 -0.916 12.5-162.5-109.8 140.4 -29.0 15.3 -4.7 54 157 A H S S+ 0 0 137 -2,-0.4 2,-0.3 1,-0.2 -1,-0.1 0.675 78.8 27.2 -91.9 -19.8 -31.6 17.9 -5.7 55 158 A D S S+ 0 0 127 2,-0.1 -1,-0.2 -37,-0.0 2,-0.1 -0.979 99.4 14.2-139.8 152.1 -34.5 16.0 -4.0 56 159 A G S S- 0 0 36 -2,-0.3 2,-0.7 -3,-0.1 -3,-0.1 -0.340 93.1 -27.2 94.9-167.3 -35.0 13.6 -1.1 57 160 A I + 0 0 149 -2,-0.1 2,-0.2 4,-0.0 -2,-0.1 -0.816 57.8 179.9 -94.4 114.3 -33.0 12.3 1.8 58 161 A A - 0 0 6 -2,-0.7 3,-0.2 1,-0.1 -41,-0.2 -0.591 40.4-110.7-100.4 171.7 -29.2 12.4 1.4 59 162 A D S S+ 0 0 34 -43,-0.3 2,-0.7 1,-0.2 -42,-0.2 0.922 111.7 41.7 -64.2 -45.6 -26.6 11.2 3.9 60 163 A I E S-c 17 0B 0 -44,-2.8 -42,-2.1 35,-0.1 2,-0.5 -0.845 74.7-168.7-111.0 97.8 -25.5 14.8 4.5 61 164 A M E -c 18 0B 27 -2,-0.7 35,-2.0 30,-0.3 2,-0.5 -0.746 9.9-160.9 -85.4 124.8 -28.3 17.3 4.7 62 165 A I E +cd 19 96B 0 -44,-2.9 -42,-2.0 -2,-0.5 -41,-0.8 -0.913 16.8 164.8-112.0 126.3 -27.0 20.8 4.5 63 166 A S E - d 0 97B 24 33,-2.4 35,-2.3 -2,-0.5 2,-0.4 -0.960 32.1-137.3-138.5 158.7 -29.0 23.8 5.7 64 167 A F E + d 0 98B 11 -2,-0.3 2,-0.3 33,-0.2 35,-0.2 -0.892 34.6 164.5-104.5 145.6 -28.8 27.4 6.7 65 168 A G E - d 0 99B 15 33,-2.4 35,-3.2 -2,-0.4 2,-0.4 -0.930 28.6-133.3-153.6 170.5 -30.8 28.6 9.7 66 169 A I - 0 0 80 -2,-0.3 33,-0.1 2,-0.3 10,-0.0 -0.997 63.4 -45.8-142.0 132.2 -31.1 31.5 12.2 67 170 A K S S+ 0 0 112 -2,-0.4 8,-1.3 2,-0.1 2,-0.5 -0.210 133.0 21.7 39.5-127.9 -31.3 31.3 16.0 68 171 A E S S+ 0 0 120 6,-0.3 -2,-0.3 1,-0.2 4,-0.1 -0.485 76.3 161.6 -60.9 113.6 -33.8 28.5 16.8 69 172 A H - 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