==== Secondary Structure Definition by the program DSSP, updated CMBI version by ElmK / April 1,2000 ==== DATE=16-AUG-2013 . REFERENCE W. KABSCH AND C.SANDER, BIOPOLYMERS 22 (1983) 2577-2637 . HEADER HYDROLASE 09-JUL-12 4G0D . COMPND 2 MOLECULE: COLLAGENASE 3; . SOURCE 2 ORGANISM_SCIENTIFIC: HOMO SAPIENS; . AUTHOR E.A.STURA,L.VERA,R.VISSE,H.NAGASE,V.DIVE . 1563 8 4 4 0 TOTAL NUMBER OF RESIDUES, NUMBER OF CHAINS, NUMBER OF SS-BRIDGES(TOTAL,INTRACHAIN,INTERCHAIN) . 70479.0 ACCESSIBLE SURFACE OF PROTEIN (ANGSTROM**2) . 933 59.7 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(J) , SAME NUMBER PER 100 RESIDUES . 78 5.0 TOTAL NUMBER OF HYDROGEN BONDS IN PARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 355 22.7 TOTAL NUMBER OF HYDROGEN BONDS IN ANTIPARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 16 1.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-5), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-4), SAME NUMBER PER 100 RESIDUES . 28 1.8 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-3), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-2), SAME NUMBER PER 100 RESIDUES . 2 0.1 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-1), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+0), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+1), SAME NUMBER PER 100 RESIDUES . 116 7.4 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+2), SAME NUMBER PER 100 RESIDUES . 167 10.7 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+3), SAME NUMBER PER 100 RESIDUES . 182 11.6 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+4), SAME NUMBER PER 100 RESIDUES . 28 1.8 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+5), SAME NUMBER PER 100 RESIDUES . 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 *** HISTOGRAMS OF *** . 0 0 0 7 0 0 0 0 0 2 4 3 1 0 0 4 0 0 0 0 0 0 0 0 0 0 0 0 0 0 RESIDUES PER ALPHA HELIX . 4 4 4 6 2 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PARALLEL BRIDGES PER LADDER . 3 18 4 5 27 12 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 ANTIPARALLEL BRIDGES PER LADDER . 6 0 8 8 0 4 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 LADDERS PER SHEET . # RESIDUE AA STRUCTURE BP1 BP2 ACC N-H-->O O-->H-N N-H-->O O-->H-N TCO KAPPA ALPHA PHI PSI X-CA Y-CA Z-CA 1 104 A Y 0 0 103 0, 0.0 2,-0.3 0, 0.0 129,-0.1 0.000 360.0 360.0 360.0 161.9 31.8 15.4 64.2 2 105 A N - 0 0 82 126,-0.4 126,-3.4 135,-0.0 2,-0.2 -0.993 360.0-152.5-138.6 142.6 34.9 17.4 65.3 3 106 A V B -A 127 0A 48 -2,-0.3 124,-0.2 124,-0.3 126,-0.1 -0.553 35.7 -84.6-101.2 171.0 38.5 16.3 66.0 4 107 A F > - 0 0 2 122,-1.3 3,-2.4 4,-0.4 -1,-0.1 -0.423 52.6 -87.2 -76.3 152.9 41.0 17.9 68.4 5 108 A P T 3 S+ 0 0 53 0, 0.0 3,-0.5 0, 0.0 -1,-0.1 -0.335 121.5 20.5 -62.1 135.1 43.1 20.9 67.4 6 109 A R T 3 S- 0 0 120 1,-0.2 3,-0.2 -3,-0.1 -2,-0.1 0.459 123.5 -96.5 79.2 3.6 46.3 19.8 65.8 7 110 A T < - 0 0 75 -3,-2.4 -1,-0.2 1,-0.2 2,-0.2 0.939 50.3-152.3 53.3 52.8 44.3 16.7 65.2 8 111 A L + 0 0 28 -3,-0.5 -4,-0.4 118,-0.1 2,-0.3 -0.390 29.3 153.8 -60.2 127.8 45.6 14.8 68.2 9 112 A K - 0 0 90 -2,-0.2 2,-0.1 -3,-0.2 117,-0.1 -0.984 48.0 -90.6-149.9 158.8 45.5 11.0 67.6 10 113 A W - 0 0 18 115,-0.4 5,-0.0 -2,-0.3 115,-0.0 -0.478 23.6-149.2 -69.7 140.1 47.3 7.8 68.8 11 114 A S S S+ 0 0 135 -2,-0.1 2,-0.3 2,-0.0 -1,-0.1 0.468 80.1 60.1 -87.0 -3.2 50.3 6.8 66.6 12 115 A K - 0 0 96 1,-0.1 -2,-0.1 2,-0.0 36,-0.0 -0.947 66.0-150.1-125.8 147.3 49.6 3.1 67.4 13 116 A M S S+ 0 0 64 -2,-0.3 36,-2.1 1,-0.1 2,-1.0 0.708 81.8 79.6 -82.7 -21.4 46.6 0.8 66.8 14 117 A N E -b 49 0B 110 34,-0.2 2,-0.3 36,-0.1 36,-0.2 -0.754 65.8-179.0 -98.0 100.5 47.3 -1.4 69.8 15 118 A L E -b 50 0B 0 34,-2.1 36,-2.8 -2,-1.0 2,-0.4 -0.765 14.4-153.4-106.2 141.8 46.0 0.4 72.9 16 119 A T E -b 51 0B 34 -2,-0.3 44,-2.5 34,-0.2 43,-0.5 -0.916 10.6-174.4-110.2 140.9 46.0 -0.7 76.5 17 120 A Y E -bc 52 60B 12 34,-2.5 36,-3.2 -2,-0.4 2,-0.3 -0.851 5.9-163.5-120.1 165.1 43.6 0.3 79.2 18 121 A R E - c 0 61B 63 42,-2.0 44,-2.8 -2,-0.3 2,-1.1 -0.960 20.2-137.4-146.3 132.9 43.5 -0.3 83.0 19 122 A I E - c 0 62B 9 34,-0.5 44,-0.2 -2,-0.3 42,-0.1 -0.779 22.6-172.0 -86.1 97.3 40.6 0.1 85.5 20 123 A V - 0 0 61 42,-2.3 2,-0.3 -2,-1.1 43,-0.2 0.884 65.1 -9.2 -60.3 -47.0 42.5 1.7 88.4 21 124 A N S S- 0 0 59 41,-0.6 2,-0.3 -3,-0.1 -1,-0.1 -0.882 72.5-121.4-142.5 178.9 39.6 1.5 90.9 22 125 A Y - 0 0 38 -2,-0.3 6,-0.1 41,-0.1 42,-0.1 -0.902 15.8-116.7-127.6 152.1 35.9 0.6 90.8 23 126 A T - 0 0 4 -2,-0.3 3,-0.4 1,-0.1 8,-0.1 -0.515 19.3-140.6 -72.5 152.1 32.5 2.2 91.6 24 127 A P S S+ 0 0 110 0, 0.0 -1,-0.1 0, 0.0 4,-0.0 0.604 92.1 82.4 -83.4 -10.7 30.7 0.4 94.4 25 128 A D S S+ 0 0 66 77,-0.1 2,-0.3 75,-0.1 78,-0.0 0.674 101.8 23.0 -68.1 -20.6 27.5 1.0 92.5 26 129 A M S S- 0 0 22 -3,-0.4 2,-0.2 79,-0.1 79,-0.1 -0.891 92.2 -96.0-137.8 164.9 28.1 -2.1 90.3 27 130 A T > - 0 0 82 -2,-0.3 4,-2.4 1,-0.1 5,-0.2 -0.539 37.1-109.6 -83.2 154.3 30.2 -5.3 90.7 28 131 A H H > S+ 0 0 103 1,-0.2 4,-2.9 2,-0.2 5,-0.2 0.871 119.0 49.6 -47.7 -45.5 33.7 -5.5 89.2 29 132 A S H > S+ 0 0 73 1,-0.2 4,-2.2 2,-0.2 -1,-0.2 0.941 110.6 49.0 -64.3 -48.0 32.6 -8.0 86.5 30 133 A E H > S+ 0 0 63 1,-0.2 4,-2.1 2,-0.2 -1,-0.2 0.856 114.3 47.2 -55.2 -39.1 29.7 -5.8 85.5 31 134 A V H X S+ 0 0 0 -4,-2.4 4,-2.1 2,-0.2 5,-0.2 0.903 110.7 50.1 -71.7 -42.0 32.0 -2.8 85.3 32 135 A E H X S+ 0 0 45 -4,-2.9 4,-1.8 -5,-0.2 -2,-0.2 0.868 111.6 51.9 -60.8 -32.8 34.7 -4.7 83.4 33 136 A K H X S+ 0 0 124 -4,-2.2 4,-2.3 -5,-0.2 -2,-0.2 0.920 107.6 49.3 -71.5 -47.8 31.9 -5.7 81.0 34 137 A A H X S+ 0 0 9 -4,-2.1 4,-2.0 1,-0.2 -2,-0.2 0.933 113.4 45.9 -57.0 -49.9 30.6 -2.1 80.4 35 138 A F H X S+ 0 0 0 -4,-2.1 4,-2.4 2,-0.2 -1,-0.2 0.880 111.8 51.8 -63.2 -39.3 34.1 -0.8 79.6 36 139 A K H X S+ 0 0 144 -4,-1.8 4,-1.1 -5,-0.2 -1,-0.2 0.905 112.0 46.7 -63.2 -40.8 34.8 -3.7 77.3 37 140 A K H X S+ 0 0 102 -4,-2.3 4,-1.0 1,-0.2 -1,-0.2 0.813 109.9 54.8 -70.1 -33.6 31.6 -3.1 75.5 38 141 A A H X S+ 0 0 0 -4,-2.0 4,-0.7 1,-0.2 3,-0.3 0.891 107.7 48.1 -69.6 -40.2 32.4 0.6 75.4 39 142 A F H X S+ 0 0 4 -4,-2.4 4,-2.6 1,-0.2 -1,-0.2 0.774 106.1 60.0 -69.7 -23.7 35.8 -0.1 73.7 40 143 A K H X S+ 0 0 109 -4,-1.1 4,-2.2 2,-0.2 -1,-0.2 0.748 91.5 65.7 -78.4 -25.9 34.1 -2.4 71.2 41 144 A V H < S+ 0 0 16 -4,-1.0 4,-0.3 -3,-0.3 -1,-0.2 0.918 114.2 33.1 -56.1 -40.5 31.8 0.4 69.9 42 145 A W H >< S+ 0 0 0 -4,-0.7 3,-1.1 -3,-0.2 4,-0.5 0.887 116.9 52.6 -87.2 -42.8 35.0 2.0 68.6 43 146 A S H >< S+ 0 0 33 -4,-2.6 3,-0.7 1,-0.2 -3,-0.2 0.814 100.4 64.9 -63.0 -29.6 37.1 -1.2 67.8 44 147 A D T 3< S+ 0 0 45 -4,-2.2 -1,-0.2 1,-0.2 122,-0.2 0.703 112.1 32.9 -66.2 -20.6 34.2 -2.5 65.6 45 148 A V T < S+ 0 0 3 -3,-1.1 -1,-0.2 -4,-0.3 -2,-0.2 0.308 120.2 48.4-120.9 6.7 34.5 0.3 63.1 46 149 A T S < S- 0 0 4 -3,-0.7 121,-0.3 -4,-0.5 120,-0.2 -0.854 86.4-107.1-136.6 170.2 38.3 1.0 63.2 47 150 A P S S+ 0 0 49 0, 0.0 -4,-0.1 0, 0.0 119,-0.1 0.436 79.5 122.8 -75.7 0.3 41.5 -1.0 63.1 48 151 A L - 0 0 5 -6,-0.3 2,-0.3 -34,-0.1 -2,-0.2 -0.345 44.0-163.8 -73.2 144.4 42.0 -0.3 66.9 49 152 A N E -b 14 0B 85 -36,-2.1 -34,-2.1 -2,-0.1 2,-0.5 -0.915 7.0-148.6-117.6 149.7 42.4 -3.1 69.5 50 153 A F E -b 15 0B 37 -2,-0.3 2,-0.5 -36,-0.2 -34,-0.2 -0.971 10.7-172.7-126.8 122.4 42.0 -2.7 73.2 51 154 A T E -b 16 0B 79 -36,-2.8 -34,-2.5 -2,-0.5 2,-0.5 -0.940 16.6-140.2-113.0 129.2 43.9 -4.8 75.8 52 155 A R E -b 17 0B 67 -2,-0.5 2,-0.3 -36,-0.2 -34,-0.2 -0.743 15.4-160.0 -90.5 127.4 43.2 -4.6 79.5 53 156 A L - 0 0 52 -36,-3.2 -34,-0.5 -2,-0.5 3,-0.1 -0.738 13.2-154.5-102.7 155.8 46.1 -4.6 82.0 54 157 A H S S+ 0 0 134 -2,-0.3 2,-0.3 1,-0.2 -1,-0.1 0.503 77.5 16.3-108.6 -7.4 45.9 -5.5 85.6 55 158 A D S S+ 0 0 122 2,-0.0 -1,-0.2 0, 0.0 2,-0.1 -0.970 99.5 26.0-154.6 160.8 48.8 -3.5 86.9 56 159 A G S S- 0 0 34 -2,-0.3 2,-0.6 -3,-0.1 -3,-0.1 -0.380 93.9 -28.6 89.1-158.9 51.1 -0.7 85.8 57 160 A I - 0 0 144 -2,-0.1 2,-0.3 4,-0.0 3,-0.1 -0.888 54.9-177.3-108.2 118.8 50.6 2.2 83.4 58 161 A A - 0 0 6 -2,-0.6 3,-0.2 1,-0.1 -41,-0.2 -0.706 38.8-114.5-103.0 162.1 48.2 1.9 80.4 59 162 A D S S+ 0 0 33 -43,-0.5 2,-0.9 -2,-0.3 -42,-0.2 0.972 112.4 42.1 -53.0 -57.1 47.5 4.4 77.7 60 163 A I E S-c 17 0B 0 -44,-2.5 -42,-2.0 35,-0.1 2,-0.6 -0.816 76.2-169.9 -98.5 103.0 43.9 4.8 79.0 61 164 A M E -c 18 0B 29 -2,-0.9 35,-1.9 30,-0.2 2,-0.4 -0.817 9.0-160.2 -88.9 123.2 43.9 4.9 82.8 62 165 A I E +cd 19 96B 0 -44,-2.8 -42,-2.3 -2,-0.6 -41,-0.6 -0.891 17.8 163.2-108.5 132.5 40.3 4.7 84.0 63 166 A S E - d 0 97B 25 33,-1.7 35,-2.5 -2,-0.4 2,-0.4 -0.981 33.4-132.6-144.8 156.6 39.2 5.8 87.5 64 167 A F E + d 0 98B 13 -2,-0.3 2,-0.3 33,-0.2 35,-0.2 -0.922 34.7 173.2-107.8 139.1 36.2 6.8 89.6 65 168 A G E - d 0 99B 10 33,-2.5 35,-2.8 -2,-0.4 36,-0.5 -0.946 24.6-144.2-145.5 159.8 36.5 10.0 91.6 66 169 A I - 0 0 84 -2,-0.3 33,-0.1 2,-0.2 -2,-0.0 -0.998 63.8 -44.4-133.6 131.2 34.4 12.3 93.8 67 170 A K S S+ 0 0 107 -2,-0.4 8,-1.7 7,-0.1 2,-0.5 -0.170 132.0 16.7 45.0-133.9 34.7 16.1 93.9 68 171 A E S S+ 0 0 93 6,-0.3 -2,-0.2 1,-0.1 4,-0.1 -0.535 77.1 164.0 -62.1 116.4 38.3 17.0 94.0 69 172 A H - 0 0 47 -2,-0.5 2,-0.3 2,-0.5 -1,-0.1 0.059 62.5 -72.2-131.3 21.9 39.9 13.7 92.9 70 173 A G S S+ 0 0 52 1,-0.5 2,-0.1 -5,-0.0 -2,-0.1 -0.438 110.5 74.7 125.4 -66.1 43.4 14.5 92.0 71 174 A D S S- 0 0 7 -2,-0.3 -1,-0.5 2,-0.2 -2,-0.5 -0.380 95.3-107.2 -65.7 163.5 43.6 16.5 88.7 72 175 A F S S+ 0 0 129 1,-0.2 -1,-0.1 -3,-0.1 -4,-0.1 0.001 99.4 92.8 -92.8 38.9 42.5 20.1 89.4 73 176 A Y - 0 0 98 -6,-0.1 -1,-0.2 11,-0.0 -2,-0.2 -0.728 66.9-164.2-124.3 79.1 39.2 19.5 87.6 74 177 A P - 0 0 63 0, 0.0 -6,-0.3 0, 0.0 2,-0.1 -0.227 15.2-118.6 -68.4 150.2 36.8 18.5 90.4 75 178 A F - 0 0 10 -8,-1.7 -7,-0.1 23,-0.1 -9,-0.1 -0.316 15.4-134.2 -82.2 176.1 33.4 16.8 90.0 76 179 A D - 0 0 70 1,-0.4 -1,-0.1 5,-0.4 -10,-0.0 0.043 37.3-106.9-127.2 20.5 30.2 18.4 91.2 77 180 A G S S+ 0 0 3 4,-0.1 -1,-0.4 3,-0.0 24,-0.2 -0.465 79.6 6.2 82.0-160.3 28.0 15.8 93.0 78 181 A P S S- 0 0 92 0, 0.0 24,-0.2 0, 0.0 3,-0.1 -0.311 120.0 -11.9 -64.5 139.1 24.8 14.3 91.6 79 182 A S S S+ 0 0 27 1,-0.2 -3,-0.2 22,-0.1 2,-0.1 -0.222 102.2 76.3 68.3-160.2 24.0 15.1 87.9 80 183 A G E S+E 1493 0B 2 1413,-2.1 1413,-1.6 -5,-0.1 2,-0.3 -0.401 98.5 28.5 64.2-132.0 25.8 17.8 86.1 81 184 A L E +E 1492 0B 49 1411,-0.2 -5,-0.4 1,-0.2 1411,-0.2 -0.467 65.3 180.0 -60.6 122.5 29.3 16.7 85.1 82 185 A L E - 0 0 0 1409,-3.3 17,-2.4 1,-0.4 2,-0.3 0.784 59.4 -36.7 -96.8 -31.3 29.2 12.9 84.6 83 186 A A E -EF1491 98B 1 1408,-0.7 1408,-1.6 15,-0.3 -1,-0.4 -0.948 54.1-141.6-174.2 172.2 32.8 12.2 83.6 84 187 A H E -EF1490 97B 7 13,-2.3 13,-2.9 -2,-0.3 2,-0.3 -0.985 9.2-161.9-152.9 162.4 35.7 13.7 81.7 85 188 A A E -EF1489 96B 0 1404,-1.8 1404,-2.4 -2,-0.3 11,-0.2 -0.997 14.8-131.3-148.3 148.8 38.5 12.6 79.4 86 189 A F - 0 0 0 9,-2.4 8,-1.0 6,-0.4 1402,-0.1 -0.764 32.3-111.3 -99.5 144.5 41.8 13.8 78.0 87 190 A P - 0 0 0 0, 0.0 7,-1.0 0, 0.0 8,-0.3 -0.102 59.2 -58.6 -62.1 168.0 42.7 13.6 74.2 88 191 A P S S+ 0 0 8 0, 0.0 6,-0.1 0, 0.0 37,-0.1 -0.173 89.4 96.8 -54.0 147.8 45.4 11.2 73.0 89 192 A G S S- 0 0 19 4,-0.3 6,-0.1 35,-0.1 -3,-0.1 -0.841 71.1 -58.2 149.3 168.4 48.8 11.7 74.6 90 193 A P S > S- 0 0 85 0, 0.0 3,-2.1 0, 0.0 2,-0.3 -0.260 101.1 -2.1 -74.4 165.2 51.2 10.5 77.4 91 194 A N T 3 S+ 0 0 129 1,-0.3 -30,-0.2 3,-0.1 -2,-0.1 -0.384 143.0 12.1 61.0-117.3 50.4 10.5 81.1 92 195 A Y T > S+ 0 0 75 -2,-0.3 3,-2.5 2,-0.1 -6,-0.4 0.759 102.6 108.4 -62.4 -32.9 46.9 12.0 81.4 93 196 A G T < S+ 0 0 1 -3,-2.1 -4,-0.3 1,-0.3 -32,-0.2 -0.255 85.9 17.1 -58.1 135.5 46.2 11.8 77.7 94 197 A G T 3 S+ 0 0 0 -8,-1.0 -1,-0.3 -7,-1.0 30,-0.2 0.283 98.2 132.9 85.5 -6.2 43.7 9.1 76.8 95 198 A D < - 0 0 9 -3,-2.5 -9,-2.4 -8,-0.3 2,-0.4 -0.376 38.5-162.9 -78.2 155.0 42.6 9.0 80.5 96 199 A A E -dF 62 85B 0 -35,-1.9 -33,-1.7 -11,-0.2 2,-0.4 -0.996 3.1-165.3-141.0 130.1 38.9 9.1 81.6 97 200 A H E -dF 63 84B 12 -13,-2.9 -13,-2.3 -2,-0.4 2,-0.4 -0.969 8.0-153.3-120.8 135.0 37.5 9.9 85.1 98 201 A F E -dF 64 83B 0 -35,-2.5 -33,-2.5 -2,-0.4 2,-0.9 -0.891 21.4-119.6-110.4 133.2 33.9 9.1 86.2 99 202 A D E > -d 65 0B 0 -17,-2.4 3,-2.1 -2,-0.4 -33,-0.2 -0.546 18.7-163.8 -71.8 105.0 32.0 11.0 88.8 100 203 A D T 3 S+ 0 0 24 -35,-2.8 -34,-0.2 -2,-0.9 -1,-0.2 0.591 81.5 80.9 -69.6 -5.0 31.1 8.4 91.5 101 204 A D T 3 S+ 0 0 49 -36,-0.5 2,-0.3 -24,-0.2 -1,-0.3 0.562 82.5 82.2 -72.9 -3.9 28.6 10.9 92.9 102 205 A E S < S- 0 0 6 -3,-2.1 2,-1.0 -24,-0.2 -77,-0.1 -0.722 91.9-119.1 -91.0 150.2 26.4 9.5 90.1 103 206 A T - 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