==== Secondary Structure Definition by the program DSSP, updated CMBI version by ElmK / April 1,2000 ==== DATE=20-JUL-2011 . REFERENCE W. KABSCH AND C.SANDER, BIOPOLYMERS 22 (1983) 2577-2637 . HEADER CYTOKINE 17-FEB-09 3GAD . COMPND 2 MOLECULE: MACROPHAGE MIGRATION INHIBITORY FACTOR-LIKE PROTE . SOURCE 2 ORGANISM_SCIENTIFIC: PLASMODIUM YOELII YOELII; . AUTHOR Y.-F.ZHOU,X.-D.SU,D.SHAO,H.WANG . 696 6 0 0 0 TOTAL NUMBER OF RESIDUES, NUMBER OF CHAINS, NUMBER OF SS-BRIDGES(TOTAL,INTRACHAIN,INTERCHAIN) . 25298.0 ACCESSIBLE SURFACE OF PROTEIN (ANGSTROM**2) . 567 81.5 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(J) , SAME NUMBER PER 100 RESIDUES . 90 12.9 TOTAL NUMBER OF HYDROGEN BONDS IN PARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 114 16.4 TOTAL NUMBER OF HYDROGEN BONDS IN ANTIPARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-5), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-4), SAME NUMBER PER 100 RESIDUES . 12 1.7 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-3), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-2), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-1), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+0), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+1), SAME NUMBER PER 100 RESIDUES . 79 11.4 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+2), SAME NUMBER PER 100 RESIDUES . 56 8.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+3), SAME NUMBER PER 100 RESIDUES . 204 29.3 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+4), SAME NUMBER PER 100 RESIDUES . 4 0.6 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+5), SAME NUMBER PER 100 RESIDUES . 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 *** HISTOGRAMS OF *** . 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 12 0 0 0 0 0 0 0 0 0 0 0 RESIDUES PER ALPHA HELIX . 0 0 0 0 0 6 6 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PARALLEL BRIDGES PER LADDER . 6 12 6 0 0 6 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 ANTIPARALLEL BRIDGES PER LADDER . 0 0 0 0 0 0 6 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 LADDERS PER SHEET . # RESIDUE AA STRUCTURE BP1 BP2 ACC N-H-->O O-->H-N N-H-->O O-->H-N TCO KAPPA ALPHA PHI PSI X-CA Y-CA Z-CA 1 2 A P 0 0 19 0, 0.0 38,-2.8 0, 0.0 2,-0.4 0.000 360.0 360.0 360.0 157.0 -14.2 -61.4 2.9 2 3 A C E -aB 39 63A 2 61,-2.7 61,-2.7 36,-0.2 2,-0.5 -0.995 360.0-169.3-127.8 132.6 -11.5 -59.2 1.4 3 4 A C E -aB 40 62A 0 36,-2.2 38,-2.0 -2,-0.4 2,-0.7 -0.914 3.3-172.3-126.8 105.5 -8.0 -60.4 0.5 4 5 A E E -aB 41 61A 14 57,-2.6 57,-2.7 -2,-0.5 2,-0.5 -0.862 3.8-166.7 -98.3 114.3 -5.5 -57.8 -0.4 5 6 A L E -aB 42 60A 0 36,-2.3 38,-3.0 -2,-0.7 2,-0.5 -0.904 4.1-174.1-100.3 128.6 -2.2 -59.1 -1.6 6 7 A I E +aB 43 59A 18 53,-3.2 53,-3.2 -2,-0.5 2,-0.3 -0.990 17.7 158.1-123.9 123.3 0.7 -56.6 -1.8 7 8 A T E -aB 44 58A 0 36,-2.2 38,-3.1 -2,-0.5 51,-0.2 -0.990 48.3-143.3-149.6 149.2 3.9 -57.8 -3.4 8 9 A N S S+ 0 0 18 49,-1.4 2,-0.6 -2,-0.3 50,-0.1 0.163 77.4 101.9 -94.7 13.6 7.1 -56.6 -5.1 9 10 A I - 0 0 29 48,-0.4 2,-0.4 46,-0.1 -2,-0.1 -0.894 63.7-149.0-100.5 123.4 7.0 -59.6 -7.4 10 11 A S + 0 0 81 -2,-0.6 -2,-0.1 34,-0.1 0, 0.0 -0.779 24.7 161.7 -92.4 132.5 5.7 -58.9 -10.9 11 12 A I - 0 0 30 -2,-0.4 5,-0.1 2,-0.0 33,-0.1 -0.963 45.0 -85.9-140.3 155.3 3.7 -61.5 -12.9 12 13 A P >> - 0 0 69 0, 0.0 4,-2.5 0, 0.0 3,-0.5 -0.214 42.8-107.5 -59.2 153.1 1.4 -61.2 -16.0 13 14 A D H 3> S+ 0 0 70 1,-0.2 4,-2.5 2,-0.2 5,-0.2 0.825 117.9 53.6 -51.6 -41.1 -2.3 -60.4 -15.4 14 15 A D H 3> S+ 0 0 116 1,-0.2 4,-1.7 2,-0.2 -1,-0.2 0.903 112.9 44.1 -64.4 -38.6 -3.5 -63.9 -16.3 15 16 A K H <> S+ 0 0 121 -3,-0.5 4,-2.2 2,-0.2 -2,-0.2 0.862 111.1 54.3 -73.7 -36.7 -1.1 -65.4 -13.8 16 17 A A H X S+ 0 0 1 -4,-2.5 4,-2.3 1,-0.2 -2,-0.2 0.944 110.5 46.3 -59.0 -48.2 -2.0 -62.8 -11.1 17 18 A Q H X S+ 0 0 35 -4,-2.5 4,-2.6 1,-0.2 -2,-0.2 0.836 108.6 55.7 -68.4 -33.5 -5.7 -63.7 -11.5 18 19 A N H X S+ 0 0 77 -4,-1.7 4,-2.1 2,-0.2 -1,-0.2 0.940 108.3 47.6 -60.2 -48.2 -5.0 -67.4 -11.3 19 20 A A H X S+ 0 0 0 -4,-2.2 4,-2.1 1,-0.2 -2,-0.2 0.939 113.3 49.3 -57.2 -47.0 -3.2 -66.9 -8.0 20 21 A L H X S+ 0 0 1 -4,-2.3 4,-2.9 1,-0.2 -1,-0.2 0.890 107.3 54.3 -58.3 -43.7 -6.1 -64.8 -6.8 21 22 A S H X S+ 0 0 54 -4,-2.6 4,-2.3 2,-0.2 -1,-0.2 0.898 109.4 48.3 -58.0 -41.9 -8.6 -67.5 -7.9 22 23 A E H X S+ 0 0 81 -4,-2.1 4,-2.2 2,-0.2 -1,-0.2 0.890 111.7 49.0 -67.0 -39.7 -6.8 -70.1 -5.9 23 24 A I H X S+ 0 0 0 -4,-2.1 4,-2.7 2,-0.2 -2,-0.2 0.914 110.1 52.2 -63.4 -43.4 -6.7 -67.8 -2.8 24 25 A E H X S+ 0 0 4 -4,-2.9 4,-2.9 2,-0.2 -2,-0.2 0.906 108.3 51.1 -58.3 -45.1 -10.4 -67.2 -3.3 25 26 A D H X S+ 0 0 100 -4,-2.3 4,-2.5 2,-0.2 -2,-0.2 0.939 111.0 47.6 -56.3 -51.2 -11.0 -71.0 -3.4 26 27 A A H X S+ 0 0 7 -4,-2.2 4,-2.6 1,-0.2 5,-0.3 0.899 113.1 48.5 -58.5 -44.6 -9.1 -71.5 -0.2 27 28 A I H X S+ 0 0 0 -4,-2.7 4,-2.4 2,-0.2 6,-0.4 0.929 110.5 51.2 -62.2 -46.2 -11.0 -68.6 1.5 28 29 A S H X>S+ 0 0 30 -4,-2.9 5,-1.0 -5,-0.2 4,-0.8 0.914 114.4 44.5 -56.6 -45.8 -14.3 -70.0 0.4 29 30 A N H <5S+ 0 0 110 -4,-2.5 3,-0.3 3,-0.2 -2,-0.2 0.961 118.2 39.3 -63.3 -57.0 -13.5 -73.4 1.8 30 31 A V H <5S+ 0 0 23 -4,-2.6 -2,-0.2 1,-0.2 -3,-0.2 0.897 123.3 38.2 -65.9 -44.9 -12.0 -72.4 5.2 31 32 A L H <5S- 0 0 20 -4,-2.4 -1,-0.2 -5,-0.3 -2,-0.2 0.550 101.5-128.3 -86.0 -7.2 -14.5 -69.6 6.0 32 33 A G T <5 + 0 0 63 -4,-0.8 -3,-0.2 -5,-0.3 -4,-0.1 0.864 63.3 136.2 60.3 39.1 -17.5 -71.4 4.5 33 34 A K < - 0 0 76 -5,-1.0 -1,-0.2 -6,-0.4 -2,-0.1 -0.951 56.7-115.3-119.6 136.0 -18.4 -68.3 2.5 34 35 A P > - 0 0 63 0, 0.0 3,-2.2 0, 0.0 4,-0.2 -0.312 26.8-113.2 -68.8 154.5 -19.4 -68.4 -1.2 35 36 A V G > S+ 0 0 51 1,-0.3 3,-2.5 2,-0.2 248,-0.6 0.804 111.7 77.3 -53.5 -31.6 -17.2 -66.8 -3.8 36 37 A A G 3 S+ 0 0 49 1,-0.3 -1,-0.3 247,-0.2 248,-0.0 0.691 97.5 44.8 -49.3 -24.7 -20.0 -64.2 -4.3 37 38 A Y G < S+ 0 0 107 -3,-2.2 2,-0.5 245,-0.1 -1,-0.3 0.405 94.2 102.1-102.6 0.1 -18.8 -62.6 -1.1 38 39 A I < - 0 0 0 -3,-2.5 245,-1.7 -4,-0.2 246,-0.6 -0.778 44.1-174.5-103.7 123.9 -15.1 -62.6 -1.8 39 40 A M E +aC 2 282A 1 -38,-2.8 -36,-2.2 -2,-0.5 2,-0.3 -0.889 12.8 174.9-103.9 140.9 -13.0 -59.7 -2.9 40 41 A S E -aC 3 281A 0 241,-2.2 241,-3.3 -2,-0.4 2,-0.3 -0.987 7.3-174.3-143.6 156.3 -9.3 -60.2 -3.9 41 42 A N E -a 4 0A 8 -38,-2.0 -36,-2.3 -2,-0.3 2,-0.5 -0.997 21.3-150.2-151.7 148.6 -6.5 -58.1 -5.3 42 43 A Y E -a 5 0A 10 -2,-0.3 2,-0.7 -38,-0.1 237,-0.3 -0.966 22.3-171.6-117.6 106.8 -2.9 -58.4 -6.5 43 44 A D E -a 6 0A 14 -38,-3.0 -36,-2.2 -2,-0.5 2,-0.6 -0.873 12.1-150.4-109.9 102.0 -1.2 -55.0 -5.9 44 45 A Y E +a 7 0A 108 -2,-0.7 2,-0.5 -38,-0.2 -36,-0.2 -0.632 19.8 175.2 -76.0 117.2 2.3 -54.8 -7.4 45 46 A Q > - 0 0 16 -38,-3.1 3,-1.9 -2,-0.6 114,-0.1 -0.886 12.3-169.3-129.2 94.7 4.4 -52.5 -5.3 46 47 A K T 3 S+ 0 0 105 -2,-0.5 -1,-0.1 1,-0.3 -38,-0.1 0.704 84.6 51.0 -64.2 -25.0 7.9 -52.5 -6.6 47 48 A N T 3 S+ 0 0 9 111,-0.3 -1,-0.3 2,-0.1 -39,-0.1 0.338 77.2 133.8 -97.5 7.5 9.4 -50.6 -3.7 48 49 A L < - 0 0 0 -3,-1.9 2,-0.3 109,-0.1 109,-0.2 -0.329 37.5-164.8 -59.1 132.1 7.9 -52.9 -1.0 49 50 A R E -H 156 0B 23 107,-3.2 107,-2.1 5,-0.1 2,-0.3 -0.935 15.8-179.9-126.3 145.7 10.5 -53.9 1.6 50 51 A F E > S+HI 155 53B 3 3,-2.5 3,-1.7 -2,-0.3 5,-0.3 -0.993 72.4 0.4-143.4 141.0 10.7 -56.6 4.3 51 52 A S T 3 S- 0 0 48 103,-1.7 101,-0.2 100,-0.4 104,-0.1 0.753 131.8 -55.0 54.9 27.8 13.5 -57.3 6.9 52 53 A G T 3 S+ 0 0 31 102,-0.6 2,-0.3 1,-0.2 -1,-0.3 0.609 118.1 91.5 85.8 13.4 15.5 -54.4 5.4 53 54 A S B < S-I 50 0B 36 -3,-1.7 -3,-2.5 2,-0.1 -1,-0.2 -0.973 70.5-135.9-141.9 159.5 15.5 -55.5 1.8 54 55 A N + 0 0 60 -2,-0.3 3,-0.1 -5,-0.2 -6,-0.1 0.122 62.0 133.3 -95.3 17.2 13.7 -55.2 -1.5 55 56 A E S S- 0 0 134 -5,-0.3 -5,-0.1 1,-0.2 -47,-0.1 -0.368 75.6 -59.8 -68.9 151.3 14.0 -59.0 -2.1 56 57 A G S S+ 0 0 31 -48,-0.1 35,-0.4 -49,-0.1 2,-0.3 0.050 74.5 160.2 -39.4 125.9 10.8 -60.9 -3.2 57 58 A Y - 0 0 0 -3,-0.1 -49,-1.4 33,-0.1 2,-0.4 -0.996 28.7-153.1-155.8 146.9 8.1 -60.6 -0.6 58 59 A C E -Bd 7 95A 0 36,-2.6 38,-2.7 -2,-0.3 2,-0.4 -0.994 4.5-170.2-128.8 129.3 4.3 -60.9 -0.1 59 60 A F E -Bd 6 96A 0 -53,-3.2 -53,-3.2 -2,-0.4 2,-0.4 -0.960 10.1-167.2-111.7 135.4 2.1 -59.1 2.4 60 61 A V E -Bd 5 97A 0 36,-2.5 38,-3.5 -2,-0.4 2,-0.6 -0.987 9.2-167.7-124.0 129.3 -1.5 -60.3 2.7 61 62 A R E -Bd 4 98A 3 -57,-2.7 -57,-2.6 -2,-0.4 2,-0.6 -0.958 9.5-171.2-113.1 113.0 -4.3 -58.5 4.5 62 63 A L E -Bd 3 99A 0 36,-3.2 38,-3.4 -2,-0.6 2,-0.5 -0.914 2.0-172.9-108.4 116.0 -7.4 -60.7 5.0 63 64 A T E +Bd 2 100A 2 -61,-2.7 -61,-2.7 -2,-0.6 2,-0.3 -0.899 10.1 168.2-109.6 130.9 -10.5 -58.9 6.4 64 65 A S E - d 0 101A 3 36,-2.6 38,-2.0 -2,-0.5 2,-1.0 -0.988 42.2-123.5-140.9 149.5 -13.6 -60.9 7.3 65 66 A I S S- 0 0 48 -2,-0.3 38,-0.1 36,-0.2 36,-0.1 -0.843 93.7 -29.1 -87.3 100.5 -16.9 -60.3 9.1 66 67 A G S S+ 0 0 31 -2,-1.0 3,-0.1 36,-0.3 36,-0.1 -0.037 104.2 93.4 71.0 175.4 -16.4 -63.1 11.7 67 68 A G + 0 0 27 1,-0.2 2,-0.8 -3,-0.1 -1,-0.1 0.474 53.3 128.4 80.4 1.1 -14.4 -66.2 10.9 68 69 A I + 0 0 28 -4,-0.1 2,-0.3 4,-0.1 -1,-0.2 -0.832 36.3 116.4 -90.1 107.0 -11.2 -64.6 12.4 69 70 A N S > S- 0 0 63 -2,-0.8 4,-2.7 -3,-0.1 5,-0.3 -0.951 77.4 -85.6-162.2 171.8 -10.0 -67.2 14.9 70 71 A R H > S+ 0 0 114 -2,-0.3 4,-2.9 1,-0.2 5,-0.2 0.931 120.8 44.4 -56.2 -52.8 -7.1 -69.5 15.6 71 72 A S H > S+ 0 0 86 1,-0.2 4,-2.4 2,-0.2 -1,-0.2 0.933 118.9 40.8 -61.5 -52.7 -8.2 -72.5 13.5 72 73 A N H > S+ 0 0 33 2,-0.2 4,-2.8 1,-0.2 5,-0.3 0.909 116.6 49.3 -62.0 -46.4 -9.3 -70.5 10.4 73 74 A N H X S+ 0 0 1 -4,-2.7 4,-2.8 2,-0.2 -2,-0.2 0.934 113.3 47.7 -59.4 -45.9 -6.3 -68.1 10.6 74 75 A S H X S+ 0 0 53 -4,-2.9 4,-1.9 -5,-0.3 -2,-0.2 0.927 113.8 46.4 -61.0 -46.8 -3.9 -71.1 10.9 75 76 A S H X S+ 0 0 31 -4,-2.4 4,-2.5 -5,-0.2 -2,-0.2 0.935 116.4 43.4 -64.0 -49.0 -5.4 -73.0 8.1 76 77 A L H X S+ 0 0 1 -4,-2.8 4,-3.1 2,-0.2 5,-0.4 0.861 109.5 57.3 -65.9 -37.5 -5.5 -70.0 5.7 77 78 A A H X S+ 0 0 0 -4,-2.8 4,-1.5 -5,-0.3 -1,-0.2 0.915 111.9 43.3 -57.0 -43.5 -2.0 -68.9 6.7 78 79 A D H X S+ 0 0 70 -4,-1.9 4,-1.8 -5,-0.2 -2,-0.2 0.938 117.9 43.5 -66.8 -48.8 -0.8 -72.3 5.6 79 80 A K H X S+ 0 0 68 -4,-2.5 4,-2.2 2,-0.2 -2,-0.2 0.907 115.4 46.3 -67.4 -45.7 -2.9 -72.5 2.4 80 81 A I H X S+ 0 0 0 -4,-3.1 4,-2.6 2,-0.2 -1,-0.2 0.885 111.2 52.3 -66.4 -39.4 -2.2 -68.9 1.2 81 82 A T H X S+ 0 0 6 -4,-1.5 4,-2.4 -5,-0.4 -1,-0.2 0.909 110.2 48.8 -64.8 -41.2 1.5 -69.1 1.9 82 83 A K H X S+ 0 0 116 -4,-1.8 4,-2.7 2,-0.2 -2,-0.2 0.926 111.1 50.4 -62.3 -46.7 1.8 -72.3 -0.2 83 84 A I H X S+ 0 0 22 -4,-2.2 4,-3.0 1,-0.2 5,-0.3 0.930 112.0 47.3 -55.8 -48.4 -0.2 -70.7 -3.1 84 85 A L H X>S+ 0 0 0 -4,-2.6 4,-2.5 1,-0.2 6,-0.8 0.895 112.8 48.1 -65.8 -40.2 2.1 -67.7 -3.1 85 86 A S H X5S+ 0 0 31 -4,-2.4 4,-1.0 4,-0.2 -1,-0.2 0.922 117.1 43.4 -64.4 -43.6 5.3 -69.7 -3.0 86 87 A N H <5S+ 0 0 134 -4,-2.7 -2,-0.2 -5,-0.2 -1,-0.2 0.934 124.7 32.3 -64.0 -48.3 4.0 -72.0 -5.8 87 88 A H H <5S+ 0 0 65 -4,-3.0 -3,-0.2 -5,-0.2 -2,-0.2 0.765 132.5 26.5 -87.0 -31.5 2.6 -69.2 -8.0 88 89 A L H <5S- 0 0 18 -4,-2.5 -3,-0.2 -5,-0.3 -2,-0.1 0.571 94.5-125.5-110.7 -14.6 4.9 -66.3 -7.3 89 90 A G << + 0 0 61 -4,-1.0 -4,-0.2 -5,-0.6 -3,-0.1 0.502 55.9 152.0 79.9 3.6 8.1 -68.1 -6.3 90 91 A V - 0 0 7 -6,-0.8 -1,-0.2 1,-0.1 -2,-0.1 -0.326 51.5-109.8 -63.4 145.9 8.3 -66.2 -3.0 91 92 A K > - 0 0 107 -35,-0.4 3,-2.6 1,-0.1 135,-0.1 -0.629 22.1-125.9 -72.8 136.5 10.1 -67.8 -0.1 92 93 A P G > S+ 0 0 57 0, 0.0 3,-1.5 0, 0.0 134,-0.8 0.858 110.9 62.9 -50.1 -36.7 7.6 -68.8 2.7 93 94 A R G 3 S+ 0 0 143 1,-0.3 -2,-0.0 133,-0.2 -3,-0.0 0.497 100.3 54.0 -71.4 -3.8 9.8 -66.8 5.1 94 95 A R G < S+ 0 0 25 -3,-2.6 -36,-2.6 131,-0.1 2,-0.5 -0.181 85.9 99.1-119.5 37.9 9.0 -63.7 3.1 95 96 A V E < -d 58 0A 0 -3,-1.5 131,-1.5 -38,-0.2 2,-0.4 -0.977 50.1-177.8-122.1 115.9 5.2 -64.1 3.3 96 97 A Y E -dE 59 225A 4 -38,-2.7 -36,-2.5 -2,-0.5 2,-0.4 -0.952 4.7-166.8-117.0 130.0 3.5 -62.0 6.0 97 98 A I E -dE 60 224A 0 127,-2.7 127,-2.4 -2,-0.4 2,-0.5 -0.971 7.5-165.5-126.4 129.6 -0.2 -62.3 6.7 98 99 A E E -dE 61 223A 0 -38,-3.5 -36,-3.2 -2,-0.4 2,-0.5 -0.935 8.6-155.1-110.3 132.5 -2.5 -60.0 8.7 99 100 A F E -d 62 0A 1 123,-3.2 2,-0.4 -2,-0.5 -36,-0.2 -0.914 12.0-170.1-107.2 130.1 -6.0 -61.1 9.7 100 101 A R E -d 63 0A 41 -38,-3.4 -36,-2.6 -2,-0.5 2,-0.4 -0.970 11.4-152.4-124.3 135.2 -8.7 -58.5 10.4 101 102 A D E -d 64 0A 100 -2,-0.4 2,-0.4 -38,-0.2 -36,-0.2 -0.845 14.4-169.1-100.9 139.2 -12.2 -58.8 11.9 102 103 A C - 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