==== Secondary Structure Definition by the program DSSP, updated CMBI version by ElmK / April 1,2000 ==== DATE=26-OCT-2012 . REFERENCE W. KABSCH AND C.SANDER, BIOPOLYMERS 22 (1983) 2577-2637 . HEADER HYDROLASE 03-SEP-12 4GWM . COMPND 2 MOLECULE: MEPRIN A SUBUNIT BETA; . SOURCE 2 ORGANISM_SCIENTIFIC: HOMO SAPIENS; . AUTHOR J.L.AROLAS,C.BRODER,T.JEFFERSON,T.GUEVARA,E.E.STERCHI,W.BODE . 1115 7 7 5 2 TOTAL NUMBER OF RESIDUES, NUMBER OF CHAINS, NUMBER OF SS-BRIDGES(TOTAL,INTRACHAIN,INTERCHAIN) . 46155.0 ACCESSIBLE SURFACE OF PROTEIN (ANGSTROM**2) . 685 61.4 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(J) , SAME NUMBER PER 100 RESIDUES . 44 3.9 TOTAL NUMBER OF HYDROGEN BONDS IN PARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 260 23.3 TOTAL NUMBER OF HYDROGEN BONDS IN ANTIPARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-5), SAME NUMBER PER 100 RESIDUES . 10 0.9 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-4), SAME NUMBER PER 100 RESIDUES . 3 0.3 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-3), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-2), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-1), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+0), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+1), SAME NUMBER PER 100 RESIDUES . 114 10.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+2), SAME NUMBER PER 100 RESIDUES . 125 11.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+3), SAME NUMBER PER 100 RESIDUES . 92 8.3 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+4), SAME NUMBER PER 100 RESIDUES . 6 0.5 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+5), SAME NUMBER PER 100 RESIDUES . 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 *** HISTOGRAMS OF *** . 0 0 0 4 0 0 0 2 0 2 0 2 0 0 0 0 2 0 0 0 0 0 0 0 0 0 0 0 0 0 RESIDUES PER ALPHA HELIX . 4 2 2 2 2 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PARALLEL BRIDGES PER LADDER . 11 2 8 7 4 2 11 2 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 ANTIPARALLEL BRIDGES PER LADDER . 2 0 2 0 2 6 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 LADDERS PER SHEET . # RESIDUE AA STRUCTURE BP1 BP2 ACC N-H-->O O-->H-N N-H-->O O-->H-N TCO KAPPA ALPHA PHI PSI X-CA Y-CA Z-CA 1 25 A E 0 0 154 0, 0.0 2,-0.1 0, 0.0 520,-0.0 0.000 360.0 360.0 360.0 20.8 -61.3 17.8 -58.8 2 26 A N - 0 0 70 519,-0.2 2,-0.6 521,-0.0 521,-0.2 -0.445 360.0-129.6 -78.9 152.2 -58.7 16.4 -56.2 3 27 A F E -a 523 0A 90 519,-3.7 521,-2.4 -2,-0.1 2,-0.7 -0.940 15.1-157.9-106.4 120.4 -56.8 13.2 -57.0 4 28 A D E > -a 524 0A 33 -2,-0.6 3,-1.6 519,-0.2 521,-0.2 -0.872 5.7-163.5-100.3 112.2 -56.7 10.5 -54.3 5 29 A V T 3 S+ 0 0 35 519,-3.1 520,-0.2 -2,-0.7 -1,-0.1 0.722 92.6 54.6 -64.7 -23.6 -53.8 8.1 -54.7 6 30 A D T > S- 0 0 18 518,-0.3 3,-1.8 -3,-0.1 -1,-0.3 0.356 105.2-131.2 -94.4 5.1 -55.5 5.7 -52.4 7 31 A G T < - 0 0 55 -3,-1.6 3,-0.1 1,-0.3 -2,-0.1 0.772 66.0 -54.3 49.2 40.8 -58.7 5.6 -54.5 8 32 A G T 3 S+ 0 0 60 1,-0.3 -1,-0.3 0, 0.0 2,-0.2 0.563 119.7 97.8 74.3 12.4 -61.2 6.2 -51.7 9 33 A M < - 0 0 94 -3,-1.8 -1,-0.3 2,-0.1 2,-0.2 -0.745 68.0-139.3-123.2 170.8 -59.8 3.2 -49.7 10 34 A D + 0 0 77 -2,-0.2 2,-0.5 -3,-0.1 -3,-0.1 -0.661 35.1 175.3-129.2 70.6 -57.4 2.5 -46.9 11 35 A Q - 0 0 62 -5,-0.2 2,-0.1 -2,-0.2 -2,-0.1 -0.685 15.0-155.0 -82.7 124.7 -55.6 -0.6 -48.2 12 36 A D >> - 0 0 19 -2,-0.5 4,-2.3 1,-0.1 3,-0.8 -0.288 42.9 -86.3 -83.9 179.5 -52.7 -2.0 -46.1 13 37 A I H 3> S+ 0 0 0 1,-0.2 4,-2.5 2,-0.2 87,-0.3 0.920 133.0 52.5 -46.9 -49.2 -49.8 -4.2 -47.6 14 38 A F H 3> S+ 0 0 2 1,-0.2 4,-1.4 2,-0.2 -1,-0.2 0.729 108.2 48.7 -67.8 -24.7 -52.1 -7.2 -47.0 15 39 A D H <> S+ 0 0 40 -3,-0.8 4,-2.0 2,-0.2 -1,-0.2 0.860 111.6 50.3 -82.1 -35.2 -55.0 -5.6 -48.9 16 40 A I H X S+ 0 0 13 -4,-2.3 4,-0.6 2,-0.2 -2,-0.2 0.972 116.4 39.6 -63.7 -55.2 -52.8 -4.7 -51.8 17 41 A N H >X>S+ 0 0 2 -4,-2.5 5,-1.4 1,-0.2 3,-0.9 0.868 114.3 53.7 -65.1 -38.0 -51.3 -8.2 -52.2 18 42 A E H ><5S+ 0 0 52 -4,-1.4 3,-1.2 1,-0.2 -1,-0.2 0.925 105.6 54.3 -60.8 -44.2 -54.7 -10.0 -51.5 19 43 A G H 3<5S+ 0 0 68 -4,-2.0 -1,-0.2 1,-0.3 -2,-0.2 0.635 110.2 47.1 -62.8 -18.5 -56.3 -7.9 -54.3 20 44 A L H <<5S- 0 0 89 -3,-0.9 -1,-0.3 -4,-0.6 -2,-0.2 0.497 106.7-127.5 -99.3 -11.8 -53.6 -9.2 -56.7 21 45 A G T <<5 + 0 0 57 -3,-1.2 -3,-0.2 -4,-0.7 -2,-0.1 0.856 52.5 153.8 65.0 43.5 -53.9 -12.9 -55.7 22 46 A L < - 0 0 52 -5,-1.4 -4,-0.1 5,-0.1 -1,-0.1 0.761 35.1-154.5 -72.4 -30.4 -50.1 -13.3 -55.0 23 47 A D + 0 0 135 -6,-0.2 2,-0.1 1,-0.2 5,-0.1 0.818 32.5 161.1 54.4 40.5 -50.6 -16.1 -52.4 24 48 A L > - 0 0 33 3,-0.3 3,-1.8 1,-0.1 2,-0.3 -0.436 54.7 -75.6 -79.8 163.9 -47.3 -15.3 -50.6 25 49 A F B >> S-G 29 0B 19 4,-1.8 4,-2.3 1,-0.3 3,-0.9 -0.443 117.4 -9.3 -63.5 122.0 -46.6 -16.6 -47.1 26 50 A E T 34 S- 0 0 3 1,-0.3 2,-0.6 -2,-0.3 -1,-0.3 0.845 126.1 -68.7 53.7 36.6 -48.6 -14.4 -44.7 27 51 A G T <4 S+ 0 0 9 -3,-1.8 -3,-0.3 1,-0.2 -1,-0.3 -0.060 131.0 31.1 81.4 -32.6 -49.4 -12.1 -47.6 28 52 A D T <4 S+ 0 0 0 -3,-0.9 74,-1.7 -2,-0.6 2,-0.5 0.325 86.1 103.6-143.4 12.9 -46.0 -10.5 -48.3 29 53 A I B < -Gh 25 102B 0 -4,-2.3 -4,-1.8 72,-0.2 2,-0.9 -0.870 52.4-152.5-104.5 126.6 -43.2 -13.0 -47.5 30 54 A R - 0 0 90 72,-2.6 74,-0.5 -2,-0.5 2,-0.2 -0.834 21.9-145.2 -94.3 103.1 -41.4 -14.8 -50.3 31 55 A L - 0 0 42 -2,-0.9 3,-0.1 72,-0.1 2,-0.1 -0.526 9.2-130.0 -72.1 143.3 -40.2 -18.1 -48.6 32 56 A D > - 0 0 54 -2,-0.2 3,-2.0 1,-0.2 2,-0.1 -0.356 46.7 -66.1 -83.0 165.7 -36.9 -19.5 -49.8 33 57 A R T 3 S- 0 0 227 1,-0.3 -1,-0.2 -2,-0.1 0, 0.0 -0.365 119.9 -9.2 -56.7 122.9 -36.5 -23.2 -50.8 34 58 A A T 3 S+ 0 0 85 1,-0.2 -1,-0.3 -3,-0.1 2,-0.2 0.576 92.6 157.1 58.9 22.1 -37.1 -25.4 -47.7 35 59 A Q < - 0 0 23 -3,-2.0 2,-0.4 1,-0.1 -1,-0.2 -0.549 44.6-122.2 -73.9 133.1 -37.2 -22.4 -45.4 36 60 A I > - 0 0 122 -2,-0.2 3,-1.2 4,-0.1 -4,-0.1 -0.669 30.2-111.3 -81.1 127.8 -39.1 -23.1 -42.1 37 61 A R T 3 S- 0 0 116 -2,-0.4 -1,-0.1 1,-0.3 100,-0.0 -0.362 95.3 -1.1 -60.0 132.8 -42.0 -20.7 -41.6 38 62 A N T 3 S+ 0 0 16 1,-0.2 -1,-0.3 129,-0.1 2,-0.2 0.816 99.2 151.3 53.3 40.3 -41.5 -18.2 -38.7 39 63 A S < - 0 0 27 -3,-1.2 -1,-0.2 1,-0.1 2,-0.1 -0.671 55.9 -87.8 -99.0 158.0 -38.1 -19.8 -37.8 40 64 A I - 0 0 138 -2,-0.2 2,-0.6 1,-0.1 -4,-0.1 -0.416 42.0-129.4 -64.0 127.1 -35.0 -18.2 -36.2 41 65 A I - 0 0 11 94,-0.3 -1,-0.1 -2,-0.1 64,-0.1 -0.774 26.8-134.3 -82.8 116.7 -32.8 -16.6 -38.9 42 66 A G > - 0 0 39 -2,-0.6 3,-1.9 1,-0.1 -1,-0.0 -0.275 22.3-102.5 -69.9 161.4 -29.3 -17.9 -38.4 43 67 A E G > S+ 0 0 182 1,-0.3 3,-1.1 2,-0.2 -1,-0.1 0.761 119.5 62.4 -52.4 -34.2 -26.1 -15.7 -38.5 44 68 A K G 3 S+ 0 0 136 1,-0.3 -1,-0.3 63,-0.0 -3,-0.0 0.745 100.1 54.6 -67.4 -24.3 -25.3 -17.0 -42.1 45 69 A Y G < S+ 0 0 77 -3,-1.9 61,-2.6 62,-0.1 2,-0.3 -0.058 88.5 106.0-103.4 35.5 -28.6 -15.4 -43.4 46 70 A R < - 0 0 39 -3,-1.1 88,-0.1 59,-0.2 89,-0.1 -0.851 66.5-124.4-114.2 148.1 -27.9 -11.9 -42.1 47 71 A W - 0 0 6 -2,-0.3 4,-0.1 87,-0.2 -2,-0.1 -0.523 33.6-103.9 -77.5 159.7 -26.8 -8.7 -44.0 48 72 A P - 0 0 76 0, 0.0 -1,-0.1 0, 0.0 176,-0.0 -0.233 44.1 -93.5 -70.7 173.0 -23.7 -6.8 -43.1 49 73 A H S S+ 0 0 88 1,-0.1 32,-2.3 31,-0.1 2,-0.5 0.773 120.0 52.6 -59.1 -29.1 -24.2 -3.5 -41.1 50 74 A T E S-i 81 0B 61 30,-0.2 32,-0.2 32,-0.0 -1,-0.1 -0.959 82.0-152.8-113.4 119.2 -24.1 -1.6 -44.5 51 75 A I E -i 82 0B 0 30,-2.9 32,-2.4 -2,-0.5 2,-0.2 -0.823 8.2-139.4 -95.2 122.1 -26.6 -3.0 -47.1 52 76 A P E -i 83 0B 7 0, 0.0 40,-2.5 0, 0.0 2,-0.3 -0.574 31.9-179.8 -76.6 138.9 -25.7 -2.5 -50.8 53 77 A Y E -ij 84 92B 20 30,-2.0 32,-2.7 -2,-0.2 2,-0.4 -0.963 27.2-155.9-139.2 161.2 -28.8 -1.6 -52.9 54 78 A V E - j 0 93B 12 38,-2.2 40,-2.9 -2,-0.3 2,-0.8 -0.998 12.3-150.3-132.2 130.0 -30.0 -0.9 -56.4 55 79 A L E - j 0 94B 10 -2,-0.4 506,-0.4 38,-0.2 40,-0.2 -0.907 31.5-139.1 -96.2 102.4 -33.2 1.2 -57.0 56 80 A E > - 0 0 47 38,-2.9 3,-1.8 -2,-0.8 8,-0.0 -0.330 13.9-116.2 -68.1 153.3 -34.2 -0.4 -60.3 57 81 A D T 3 S+ 0 0 79 1,-0.3 -1,-0.1 504,-0.1 38,-0.1 0.646 109.2 71.5 -62.4 -20.5 -35.6 1.8 -63.2 58 82 A S T 3 S+ 0 0 61 2,-0.1 -1,-0.3 0, 0.0 37,-0.1 0.736 77.3 104.5 -69.3 -21.8 -39.0 0.0 -63.1 59 83 A L S < S- 0 0 7 -3,-1.8 58,-0.1 35,-0.2 2,-0.1 -0.299 81.8-110.9 -61.3 140.5 -39.7 1.9 -59.7 60 84 A E > - 0 0 68 56,-0.3 4,-1.9 1,-0.1 3,-0.1 -0.398 27.8-116.2 -63.8 147.5 -42.1 4.8 -59.8 61 85 A M H > S+ 0 0 19 1,-0.2 4,-1.9 2,-0.2 5,-0.2 0.918 117.1 51.0 -54.1 -44.1 -40.3 8.2 -59.3 62 86 A N H > S+ 0 0 16 1,-0.2 4,-2.2 2,-0.2 -1,-0.2 0.881 107.8 53.9 -61.8 -38.2 -42.3 8.7 -56.0 63 87 A A H > S+ 0 0 2 2,-0.2 4,-2.6 1,-0.2 -1,-0.2 0.894 104.4 54.0 -63.4 -41.2 -41.3 5.2 -54.8 64 88 A K H X S+ 0 0 9 -4,-1.9 4,-2.0 1,-0.2 -2,-0.2 0.942 112.0 44.8 -60.2 -46.0 -37.6 6.0 -55.4 65 89 A G H X S+ 0 0 3 -4,-1.9 4,-1.8 1,-0.2 -1,-0.2 0.871 112.8 50.8 -63.5 -41.2 -37.8 9.1 -53.2 66 90 A V H X S+ 0 0 3 -4,-2.2 4,-2.2 2,-0.2 -1,-0.2 0.889 108.8 51.8 -63.7 -42.6 -39.9 7.3 -50.5 67 91 A I H X S+ 0 0 0 -4,-2.6 4,-2.0 2,-0.2 -2,-0.2 0.936 109.4 48.9 -62.2 -45.5 -37.3 4.4 -50.3 68 92 A L H X S+ 0 0 45 -4,-2.0 4,-1.4 1,-0.2 -1,-0.2 0.860 109.9 53.1 -63.8 -32.0 -34.4 6.9 -49.8 69 93 A N H X S+ 0 0 32 -4,-1.8 4,-1.3 2,-0.2 -1,-0.2 0.887 106.3 53.2 -65.4 -40.1 -36.6 8.6 -47.1 70 94 A A H X S+ 0 0 0 -4,-2.2 4,-1.8 1,-0.2 3,-0.3 0.895 104.9 54.8 -61.8 -39.9 -36.9 5.1 -45.4 71 95 A F H X S+ 0 0 0 -4,-2.0 4,-2.4 1,-0.2 -1,-0.2 0.874 103.1 56.2 -61.7 -37.3 -33.1 4.8 -45.4 72 96 A E H X S+ 0 0 57 -4,-1.4 4,-1.5 2,-0.2 -1,-0.2 0.858 104.6 52.6 -64.1 -33.9 -32.9 8.1 -43.6 73 97 A R H X S+ 0 0 23 -4,-1.3 4,-2.2 -3,-0.3 -1,-0.2 0.911 109.6 48.2 -67.1 -41.3 -35.1 6.6 -40.8 74 98 A Y H X S+ 0 0 0 -4,-1.8 4,-2.0 1,-0.2 6,-0.4 0.907 110.7 53.0 -61.4 -40.3 -32.8 3.6 -40.5 75 99 A R H < S+ 0 0 32 -4,-2.4 156,-2.3 1,-0.2 155,-1.1 0.778 113.3 42.9 -69.0 -26.2 -29.8 6.1 -40.3 76 100 A L H < S+ 0 0 22 -4,-1.5 -1,-0.2 155,-0.2 156,-0.2 0.872 125.3 29.5 -84.1 -39.0 -31.4 8.0 -37.5 77 101 A K H < S+ 0 0 15 -4,-2.2 2,-0.3 -5,-0.1 -2,-0.2 0.518 120.8 31.7-105.0 -7.0 -32.7 5.2 -35.3 78 102 A T S < S- 0 0 1 -4,-2.0 152,-0.7 2,-0.2 154,-0.1 -0.886 73.1-116.1-139.1 174.5 -30.2 2.3 -36.0 79 103 A a S S+ 0 0 33 -2,-0.3 2,-0.3 150,-0.2 -4,-0.1 0.297 81.5 113.5 -97.2 14.6 -26.6 1.5 -36.8 80 104 A I - 0 0 2 -6,-0.4 2,-0.3 -30,-0.1 -2,-0.2 -0.658 45.9-177.9 -81.8 134.9 -27.9 -0.0 -40.2 81 105 A D E -i 50 0B 40 -32,-2.3 -30,-2.9 -2,-0.3 2,-0.4 -0.839 16.7-142.3-130.2 166.0 -26.8 1.9 -43.3 82 106 A F E -i 51 0B 4 -2,-0.3 -32,-0.0 -32,-0.2 -33,-0.0 -0.987 16.2-178.5-130.3 139.8 -27.4 1.6 -47.0 83 107 A K E -i 52 0B 68 -32,-2.4 -30,-2.0 -2,-0.4 3,-0.1 -0.994 37.8 -93.7-139.2 145.9 -25.1 2.2 -49.8 84 108 A P E -i 53 0B 66 0, 0.0 -30,-0.2 0, 0.0 478,-0.0 -0.334 56.9 -95.4 -54.9 136.9 -25.4 2.0 -53.6 85 109 A W + 0 0 73 -32,-2.7 6,-0.1 7,-0.1 -34,-0.0 -0.281 44.2 176.2 -56.9 130.4 -24.3 -1.4 -54.9 86 110 A A S S- 0 0 41 -3,-0.1 -1,-0.1 0, 0.0 476,-0.0 0.012 71.2 -63.0-127.3 26.3 -20.6 -1.3 -56.0 87 111 A G S S+ 0 0 52 1,-0.2 -2,-0.1 476,-0.1 4,-0.1 0.448 79.4 156.4 107.7 3.2 -20.1 -5.0 -56.9 88 112 A E - 0 0 68 1,-0.1 -1,-0.2 2,-0.1 3,-0.0 -0.180 52.9-115.5 -55.4 152.4 -20.6 -6.7 -53.5 89 113 A T S S+ 0 0 122 1,-0.1 2,-0.4 22,-0.0 -1,-0.1 0.900 98.8 34.9 -64.8 -40.9 -21.6 -10.4 -53.8 90 114 A N S S+ 0 0 24 19,-0.1 2,-0.3 21,-0.1 21,-0.2 -0.930 71.1 154.7-118.6 138.4 -25.1 -9.9 -52.3 91 115 A Y E - 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