==== Secondary Structure Definition by the program DSSP, updated CMBI version by ElmK / April 1,2000 ==== DATE=31-DEC-2009 . REFERENCE W. KABSCH AND C.SANDER, BIOPOLYMERS 22 (1983) 2577-2637 . HEADER TRANSFERASE 18-MAY-06 2H29 . COMPND 2 MOLECULE: PROBABLE NICOTINATE-NUCLEOTIDE . SOURCE 2 ORGANISM_SCIENTIFIC: STAPHYLOCOCCUS AUREUS; . AUTHOR S.HAN . 376 2 0 0 0 TOTAL NUMBER OF RESIDUES, NUMBER OF CHAINS, NUMBER OF SS-BRIDGES(TOTAL,INTRACHAIN,INTERCHAIN) . 18235.0 ACCESSIBLE SURFACE OF PROTEIN (ANGSTROM**2) . 250 66.5 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(J) , SAME NUMBER PER 100 RESIDUES . 61 16.2 TOTAL NUMBER OF HYDROGEN BONDS IN PARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 2 0.5 TOTAL NUMBER OF HYDROGEN BONDS IN ANTIPARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-5), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-4), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-3), SAME NUMBER PER 100 RESIDUES . 2 0.5 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-2), SAME NUMBER PER 100 RESIDUES . 2 0.5 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-1), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+0), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+1), SAME NUMBER PER 100 RESIDUES . 22 5.9 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+2), SAME NUMBER PER 100 RESIDUES . 38 10.1 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+3), SAME NUMBER PER 100 RESIDUES . 116 30.9 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+4), SAME NUMBER PER 100 RESIDUES . 6 1.6 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+5), SAME NUMBER PER 100 RESIDUES . 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 *** HISTOGRAMS OF *** . 0 0 0 2 2 0 2 0 1 4 1 0 0 4 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 RESIDUES PER ALPHA HELIX . 4 2 2 0 3 3 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PARALLEL BRIDGES PER LADDER . 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 ANTIPARALLEL BRIDGES PER LADDER . 0 0 0 0 2 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 LADDERS PER SHEET . # RESIDUE AA STRUCTURE BP1 BP2 ACC N-H-->O O-->H-N N-H-->O O-->H-N TCO KAPPA ALPHA PHI PSI X-CA Y-CA Z-CA 1 1 A M 0 0 188 0, 0.0 2,-0.2 0, 0.0 98,-0.2 0.000 360.0 360.0 360.0 -67.7 48.2 27.4 -7.7 2 2 A K E -a 99 0A 66 96,-1.0 98,-2.7 29,-0.1 2,-0.6 -0.606 360.0-125.9 -72.5 142.4 48.7 26.9 -4.0 3 3 A K E -a 100 0A 63 -2,-0.2 30,-1.8 96,-0.2 29,-1.8 -0.838 25.9-173.3 -94.7 121.2 45.4 26.1 -2.2 4 4 A I E -ab 101 33A 0 96,-2.9 98,-2.5 -2,-0.6 2,-0.4 -0.977 4.0-164.7-115.8 126.4 44.5 28.3 0.8 5 5 A V E -ab 102 34A 0 28,-2.5 30,-3.4 -2,-0.5 2,-0.5 -0.906 1.9-168.5-106.2 132.1 41.5 27.4 3.0 6 6 A L E -ab 103 35A 0 96,-2.9 98,-2.2 -2,-0.4 2,-0.6 -0.989 5.2-161.3-117.8 135.9 40.0 29.9 5.4 7 7 A Y E -ab 104 36A 20 28,-3.6 30,-3.1 -2,-0.5 2,-0.2 -0.924 16.2-157.4-118.2 106.0 37.5 28.8 7.9 8 8 A G E + b 0 37A 21 96,-0.7 2,-0.3 -2,-0.6 3,-0.1 -0.528 26.3 124.6 -91.7 148.7 35.5 31.8 9.2 9 9 A G E - b 0 38A 18 28,-1.9 30,-2.3 -2,-0.2 5,-0.1 -0.949 63.9-110.7 178.1 163.6 33.6 32.0 12.5 10 10 A Q - 0 0 48 -2,-0.3 4,-0.1 28,-0.2 -1,-0.1 0.823 43.7-164.6 -70.9 -36.7 32.8 33.8 15.8 11 11 A F > + 0 0 3 2,-0.2 3,-0.9 1,-0.1 49,-0.1 0.946 16.1 173.1 46.0 61.8 34.3 30.7 17.5 12 12 A N T 3 S- 0 0 37 1,-0.9 148,-0.2 47,-0.0 -1,-0.1 -0.509 87.2 -3.6-132.8 68.7 32.8 31.7 20.9 13 13 A P T 3 S- 0 0 0 0, 0.0 -1,-0.9 0, 0.0 -2,-0.2 0.486 90.0-138.0 -93.9 164.3 33.8 29.1 21.8 14 14 A I < - 0 0 1 -3,-0.9 2,-0.3 42,-0.2 141,-0.1 -0.427 38.3-179.4 -60.0 146.8 35.4 26.9 19.2 15 15 A H > - 0 0 4 158,-0.1 4,-1.5 -2,-0.1 158,-0.2 -0.843 42.0 -79.2-147.9-178.4 34.2 23.3 19.8 16 16 A T H > S+ 0 0 10 156,-2.9 4,-1.7 -2,-0.3 5,-0.1 0.811 122.7 55.2 -64.3 -35.0 34.3 19.7 18.6 17 17 A A H > S+ 0 0 12 155,-0.4 4,-2.5 1,-0.2 -1,-0.2 0.935 106.2 52.3 -65.0 -38.6 32.0 20.2 15.7 18 18 A H H > S+ 0 0 28 1,-0.2 4,-2.5 2,-0.2 -2,-0.2 0.881 109.0 51.5 -59.6 -36.7 34.3 23.0 14.4 19 19 A M H X S+ 0 0 1 -4,-1.5 4,-2.6 2,-0.2 -1,-0.2 0.844 107.3 51.5 -68.8 -37.1 37.2 20.6 14.6 20 20 A I H X S+ 0 0 1 -4,-1.7 4,-2.5 2,-0.2 5,-0.2 0.929 111.6 48.4 -65.5 -44.8 35.3 17.9 12.7 21 21 A V H X S+ 0 0 10 -4,-2.5 4,-2.3 2,-0.2 5,-0.3 0.946 113.9 45.1 -57.5 -53.1 34.6 20.5 10.0 22 22 A A H X S+ 0 0 0 -4,-2.5 4,-3.0 1,-0.2 5,-0.2 0.951 114.6 48.9 -58.9 -48.9 38.2 21.7 9.8 23 23 A S H X S+ 0 0 0 -4,-2.6 4,-2.4 1,-0.2 -1,-0.2 0.899 112.5 46.1 -60.7 -47.6 39.6 18.2 9.8 24 24 A E H X S+ 0 0 1 -4,-2.5 4,-2.3 2,-0.2 -1,-0.2 0.895 115.8 45.2 -64.1 -43.2 37.4 16.8 7.1 25 25 A V H X S+ 0 0 0 -4,-2.3 4,-3.2 -5,-0.2 5,-0.4 0.939 113.7 50.7 -65.0 -44.8 37.9 19.8 4.7 26 26 A F H X S+ 0 0 54 -4,-3.0 4,-1.9 -5,-0.3 -2,-0.2 0.904 114.4 43.0 -61.4 -44.7 41.7 19.8 5.4 27 27 A H H < S+ 0 0 78 -4,-2.4 -1,-0.2 -5,-0.2 -2,-0.2 0.838 118.2 44.5 -72.1 -37.4 41.9 16.0 4.6 28 28 A E H < S+ 0 0 90 -4,-2.3 -2,-0.2 -5,-0.2 -3,-0.2 0.935 124.8 31.8 -70.0 -48.4 39.6 16.2 1.5 29 29 A L H < S- 0 0 17 -4,-3.2 -2,-0.2 -5,-0.2 -3,-0.2 0.681 85.0-154.2 -87.2 -21.4 41.1 19.3 -0.0 30 30 A Q < - 0 0 162 -4,-1.9 -27,-0.2 -5,-0.4 -3,-0.1 0.918 24.7-169.9 48.3 56.2 44.7 18.9 1.1 31 31 A P - 0 0 12 0, 0.0 -27,-0.2 0, 0.0 3,-0.1 -0.277 33.1-127.0 -77.7 164.5 45.5 22.6 1.0 32 32 A D S S+ 0 0 110 -29,-1.8 2,-0.3 1,-0.2 -28,-0.2 0.840 97.3 19.4 -71.6 -33.8 48.9 24.2 1.3 33 33 A E E -b 4 0A 52 -30,-1.8 -28,-2.5 36,-0.0 2,-0.4 -0.995 63.8-165.4-139.8 142.5 47.6 26.5 4.1 34 34 A F E -b 5 0A 3 -2,-0.3 37,-1.7 -30,-0.2 2,-0.5 -0.969 12.3-168.0-130.2 118.1 44.5 26.3 6.4 35 35 A Y E -bc 6 71A 37 -30,-3.4 -28,-3.6 -2,-0.4 2,-0.5 -0.908 19.5-148.4-113.3 128.2 43.4 29.4 8.4 36 36 A F E -bc 7 72A 0 35,-2.8 37,-2.6 -2,-0.5 -28,-0.2 -0.785 23.9-157.8 -75.9 128.8 40.9 29.6 11.2 37 37 A L E -b 8 0A 1 -30,-3.1 -28,-1.9 -2,-0.5 2,-0.4 -0.828 3.7-161.6-119.2 88.1 39.5 33.1 10.7 38 38 A P E -b 9 0A 0 0, 0.0 39,-0.3 0, 0.0 38,-0.2 -0.576 16.0-133.9 -72.7 129.2 38.0 34.5 13.9 39 39 A S - 0 0 4 -30,-2.3 3,-0.2 -2,-0.4 38,-0.1 -0.421 8.3-148.7 -72.3 150.1 35.6 37.5 13.3 40 40 A F S S- 0 0 56 1,-0.3 2,-0.4 41,-0.1 -1,-0.2 0.912 97.7 -5.5 -74.9 -58.6 35.8 40.7 15.3 41 41 A M B S-f 82 0B 30 40,-0.6 42,-3.3 39,-0.1 -1,-0.3 -0.941 79.8-140.1-135.5 109.9 32.0 41.0 14.8 42 42 A S > - 0 0 24 -2,-0.4 3,-1.0 -3,-0.2 42,-0.1 -0.568 30.1-113.9 -67.2 125.6 30.4 38.4 12.5 43 43 A P T 3 S+ 0 0 31 0, 0.0 3,-0.1 0, 0.0 -1,-0.1 -0.303 98.1 28.2 -58.9 146.2 27.7 40.1 10.3 44 44 A L T 3 S+ 0 0 95 1,-0.3 2,-0.3 -3,-0.0 -2,-0.0 0.654 105.0 91.1 75.4 18.5 24.1 39.1 11.0 45 45 A K S < S- 0 0 90 -3,-1.0 -1,-0.3 0, 0.0 2,-0.2 -0.931 76.5-114.6-133.0 158.0 24.8 38.1 14.6 46 46 A K - 0 0 203 -2,-0.3 2,-0.3 -3,-0.1 -4,-0.1 -0.553 31.2-160.3 -87.7 159.0 24.5 40.2 17.8 47 47 A H - 0 0 98 -2,-0.2 4,-0.1 -6,-0.1 2,-0.1 -0.947 19.7-107.5-134.6 158.7 27.5 41.0 19.9 48 48 A H - 0 0 149 -2,-0.3 0, 0.0 2,-0.2 0, 0.0 -0.325 37.5-109.3 -69.5 164.6 28.3 42.2 23.5 49 49 A D S S+ 0 0 169 1,-0.1 2,-0.5 -2,-0.1 -1,-0.1 0.941 111.9 51.5 -59.9 -49.5 29.5 45.7 24.1 50 50 A F S S- 0 0 160 -3,-0.0 2,-0.7 2,-0.0 -2,-0.2 -0.808 82.7-148.5 -89.5 127.5 33.0 44.4 24.9 51 51 A I - 0 0 29 -2,-0.5 2,-0.3 -4,-0.1 -10,-0.1 -0.866 11.5-142.6 -98.3 112.1 34.4 42.0 22.2 52 52 A D + 0 0 29 -2,-0.7 -4,-0.0 1,-0.2 -2,-0.0 -0.582 33.2 158.2 -72.3 135.5 36.7 39.4 23.6 53 53 A V > + 0 0 36 -2,-0.3 4,-2.4 3,-0.1 5,-0.2 0.684 65.1 58.3-116.5 -50.3 39.6 38.5 21.5 54 54 A Q H > S+ 0 0 123 1,-0.2 4,-2.2 2,-0.2 5,-0.2 0.876 107.5 44.1 -52.3 -53.2 42.2 37.0 23.9 55 55 A H H > S+ 0 0 39 2,-0.2 4,-2.4 1,-0.2 -1,-0.2 0.923 115.0 49.2 -62.8 -45.4 40.2 34.0 25.2 56 56 A R H > S+ 0 0 13 2,-0.2 4,-2.5 1,-0.2 -2,-0.2 0.936 112.5 46.7 -61.4 -50.4 38.8 33.1 21.7 57 57 A L H X S+ 0 0 36 -4,-2.4 4,-2.6 2,-0.2 -1,-0.2 0.877 111.4 52.3 -56.1 -46.2 42.3 33.1 20.1 58 58 A T H X S+ 0 0 52 -4,-2.2 4,-1.6 -5,-0.2 -2,-0.2 0.937 111.9 45.9 -55.3 -49.0 43.7 31.1 23.0 59 59 A M H X S+ 0 0 0 -4,-2.4 4,-1.3 2,-0.2 -2,-0.2 0.883 111.5 51.2 -66.6 -38.4 41.0 28.5 22.6 60 60 A I H X S+ 0 0 0 -4,-2.5 4,-2.3 1,-0.2 3,-0.4 0.923 107.6 53.0 -62.8 -44.7 41.4 28.4 18.7 61 61 A Q H X S+ 0 0 81 -4,-2.6 4,-2.4 1,-0.2 -1,-0.2 0.878 104.8 56.7 -56.6 -35.0 45.2 27.8 19.2 62 62 A M H X S+ 0 0 39 -4,-1.6 4,-1.7 2,-0.2 -1,-0.2 0.869 107.0 48.3 -65.4 -32.9 44.2 24.9 21.5 63 63 A I H X S+ 0 0 0 -4,-1.3 4,-2.8 -3,-0.4 5,-0.2 0.923 109.1 51.9 -70.7 -45.1 42.2 23.3 18.7 64 64 A I H X>S+ 0 0 15 -4,-2.3 4,-2.1 1,-0.2 5,-0.6 0.891 108.2 54.0 -55.4 -42.9 45.2 23.8 16.3 65 65 A D H <5S+ 0 0 101 -4,-2.4 -1,-0.2 1,-0.2 -2,-0.2 0.918 111.9 42.8 -57.8 -47.1 47.4 22.1 18.9 66 66 A E H <5S+ 0 0 55 -4,-1.7 -2,-0.2 1,-0.2 -1,-0.2 0.917 115.7 48.4 -67.4 -41.3 45.1 19.0 19.0 67 67 A L H <5S- 0 0 8 -4,-2.8 -2,-0.2 2,-0.2 -1,-0.2 0.779 95.0-137.2 -72.1 -32.5 44.6 18.9 15.2 68 68 A G T <5S+ 0 0 68 -4,-2.1 2,-0.3 1,-0.3 -3,-0.1 0.623 75.6 67.3 80.9 12.2 48.2 19.1 14.2 69 69 A F < + 0 0 23 -5,-0.6 -1,-0.3 -36,-0.0 -2,-0.2 -0.967 64.9 59.8-149.0 167.3 47.5 21.6 11.3 70 70 A G - 0 0 14 -2,-0.3 2,-0.3 -3,-0.1 -35,-0.2 -0.112 45.5-134.5 103.0 163.1 46.2 25.1 10.8 71 71 A D E -c 35 0A 90 -37,-1.7 -35,-2.8 -2,-0.1 2,-0.4 -0.939 23.7-111.6-146.5 159.3 47.0 28.7 11.6 72 72 A I E -c 36 0A 34 -2,-0.3 2,-0.6 -37,-0.2 -35,-0.2 -0.847 19.8-150.7 -93.7 134.7 45.0 31.7 12.9 73 73 A C - 0 0 7 -37,-2.6 3,-0.4 -2,-0.4 4,-0.2 -0.953 9.9-171.4-103.0 116.1 44.5 34.6 10.5 74 74 A D > + 0 0 61 -2,-0.6 4,-2.6 1,-0.2 3,-0.3 0.333 50.4 111.8 -90.7 4.6 44.1 37.7 12.6 75 75 A D H > S+ 0 0 38 1,-0.2 4,-2.3 2,-0.2 -1,-0.2 0.855 80.9 44.1 -46.4 -49.6 43.1 40.1 9.8 76 76 A E H > S+ 0 0 5 -3,-0.4 4,-1.9 -38,-0.2 -1,-0.2 0.856 113.1 51.3 -71.3 -34.2 39.5 40.6 11.1 77 77 A I H 4 S+ 0 0 30 -3,-0.3 -2,-0.2 -39,-0.3 -1,-0.2 0.926 113.2 46.1 -62.4 -46.7 40.7 41.0 14.7 78 78 A K H < S+ 0 0 152 -4,-2.6 -2,-0.2 1,-0.2 -1,-0.2 0.880 115.1 47.1 -62.0 -42.0 43.2 43.7 13.6 79 79 A R H < S- 0 0 100 -4,-2.3 -1,-0.2 -5,-0.2 -2,-0.2 0.829 90.9-169.8 -71.6 -36.2 40.7 45.5 11.4 80 80 A G < + 0 0 17 -4,-1.9 -40,-0.1 -5,-0.2 -39,-0.1 -0.124 40.6 74.7 67.3-170.8 37.8 45.6 13.9 81 81 A G S S- 0 0 39 -41,-0.1 -40,-0.6 -4,-0.1 2,-0.4 -0.207 102.7 -13.2 68.1-164.7 34.3 46.7 13.1 82 82 A Q B S-f 41 0B 66 -42,-0.1 2,-1.7 1,-0.1 -40,-0.2 -0.601 70.5-133.1 -67.8 125.9 31.8 44.4 11.3 83 83 A S - 0 0 2 -42,-3.3 2,-0.3 -2,-0.4 -1,-0.1 -0.536 27.5-175.4 -90.5 75.7 33.8 41.6 9.7 84 84 A Y > - 0 0 77 -2,-1.7 4,-1.8 1,-0.2 3,-0.3 -0.552 22.1-143.2 -71.9 130.2 32.5 41.5 6.1 85 85 A T H > S+ 0 0 19 -2,-0.3 4,-3.0 1,-0.2 5,-0.3 0.868 96.4 61.6 -67.3 -36.1 34.0 38.6 4.3 86 86 A Y H > S+ 0 0 38 31,-0.4 4,-2.6 1,-0.2 -1,-0.2 0.929 106.8 45.4 -52.5 -47.3 34.3 40.5 1.0 87 87 A D H > S+ 0 0 54 -3,-0.3 4,-2.2 1,-0.2 5,-0.2 0.918 113.3 49.9 -66.7 -45.0 36.6 43.1 2.6 88 88 A T H X S+ 0 0 0 -4,-1.8 4,-2.6 1,-0.2 -1,-0.2 0.919 113.9 44.7 -54.9 -48.4 38.7 40.5 4.3 89 89 A I H X S+ 0 0 0 -4,-3.0 4,-3.2 2,-0.2 5,-0.3 0.903 109.4 54.7 -70.3 -41.5 39.1 38.4 1.1 90 90 A K H X S+ 0 0 89 -4,-2.6 4,-1.4 -5,-0.3 -1,-0.2 0.954 115.9 40.1 -53.6 -50.6 39.9 41.5 -1.1 91 91 A A H X S+ 0 0 45 -4,-2.2 4,-0.7 -5,-0.2 -2,-0.2 0.894 115.7 51.3 -68.1 -39.9 42.7 42.5 1.3 92 92 A F H >X S+ 0 0 6 -4,-2.6 3,-0.8 -5,-0.2 4,-0.6 0.921 107.5 51.0 -63.7 -46.3 43.9 38.9 1.8 93 93 A K H >< S+ 0 0 47 -4,-3.2 3,-1.0 1,-0.2 6,-0.2 0.878 104.0 57.2 -64.1 -36.1 44.2 38.0 -1.9 94 94 A E H 3< S+ 0 0 107 -4,-1.4 3,-0.3 -5,-0.3 -1,-0.2 0.755 111.2 45.7 -66.4 -19.1 46.3 41.1 -2.7 95 95 A Q H << S+ 0 0 153 -3,-0.8 -1,-0.3 -4,-0.7 -2,-0.2 0.520 123.1 35.5 -93.0 -9.3 48.7 39.7 -0.1 96 96 A H S X< S+ 0 0 44 -3,-1.0 3,-1.3 -4,-0.6 -1,-0.2 -0.367 77.3 167.2-142.4 50.8 48.5 36.1 -1.5 97 97 A K T 3 + 0 0 113 -3,-0.3 3,-0.1 1,-0.2 -3,-0.1 -0.324 68.3 38.2 -62.8 152.6 48.2 36.6 -5.3 98 98 A D T 3 S+ 0 0 145 1,-0.3 -96,-1.0 -97,-0.1 2,-0.4 0.497 95.5 111.1 78.6 2.6 48.7 33.3 -7.2 99 99 A S E < -a 2 0A 19 -3,-1.3 2,-0.5 -6,-0.2 -1,-0.3 -0.916 67.4-131.6-108.9 137.5 46.8 31.4 -4.4 100 100 A E E -a 3 0A 102 -98,-2.7 -96,-2.9 -2,-0.4 2,-0.4 -0.781 27.8-142.7 -83.2 126.8 43.4 29.8 -4.9 101 101 A L E -a 4 0A 8 -2,-0.5 26,-2.2 24,-0.4 2,-0.4 -0.776 17.4-172.9-102.7 131.7 41.2 30.8 -2.1 102 102 A Y E -ad 5 127A 26 -98,-2.5 -96,-2.9 -2,-0.4 2,-0.5 -0.974 6.1-161.1-122.6 133.9 38.6 28.5 -0.4 103 103 A F E -ad 6 128A 6 24,-2.3 26,-2.8 -2,-0.4 2,-0.4 -0.960 10.9-146.6-124.1 118.3 36.1 29.6 2.2 104 104 A V E +ad 7 129A 1 -98,-2.2 -96,-0.7 -2,-0.5 2,-0.3 -0.678 26.1 161.9 -89.1 126.7 34.4 27.0 4.4 105 105 A I E - 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