==== Secondary Structure Definition by the program DSSP, updated CMBI version by ElmK / April 1,2000 ==== DATE=22-MAR-2013 . REFERENCE W. KABSCH AND C.SANDER, BIOPOLYMERS 22 (1983) 2577-2637 . HEADER OXIDOREDUCTASE/OXIDOREDUCTASE INHIBITOR 24-SEP-12 4H95 . COMPND 2 MOLECULE: DIHYDROFOLATE REDUCTASE; . SOURCE 2 ORGANISM_SCIENTIFIC: CANDIDA ALBICANS; . AUTHOR J.L.PAULSEN,A.C.ANDERSON . 378 2 0 0 0 TOTAL NUMBER OF RESIDUES, NUMBER OF CHAINS, NUMBER OF SS-BRIDGES(TOTAL,INTRACHAIN,INTERCHAIN) . 20533.0 ACCESSIBLE SURFACE OF PROTEIN (ANGSTROM**2) . 261 69.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(J) , SAME NUMBER PER 100 RESIDUES . 63 16.7 TOTAL NUMBER OF HYDROGEN BONDS IN PARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 57 15.1 TOTAL NUMBER OF HYDROGEN BONDS IN ANTIPARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 2 0.5 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-5), SAME NUMBER PER 100 RESIDUES . 2 0.5 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-4), SAME NUMBER PER 100 RESIDUES . 6 1.6 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-3), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-2), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-1), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+0), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+1), SAME NUMBER PER 100 RESIDUES . 39 10.3 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+2), SAME NUMBER PER 100 RESIDUES . 35 9.3 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+3), SAME NUMBER PER 100 RESIDUES . 64 16.9 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+4), SAME NUMBER PER 100 RESIDUES . 2 0.5 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+5), SAME NUMBER PER 100 RESIDUES . 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 *** HISTOGRAMS OF *** . 0 0 0 0 1 2 3 2 0 0 0 2 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 RESIDUES PER ALPHA HELIX . 0 0 0 6 1 1 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PARALLEL BRIDGES PER LADDER . 4 4 3 1 0 0 0 2 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 ANTIPARALLEL BRIDGES PER LADDER . 0 0 0 0 0 0 0 0 0 0 2 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 LADDERS PER SHEET . # RESIDUE AA STRUCTURE BP1 BP2 ACC N-H-->O O-->H-N N-H-->O O-->H-N TCO KAPPA ALPHA PHI PSI X-CA Y-CA Z-CA 1 4 A P 0 0 118 0, 0.0 2,-0.4 0, 0.0 104,-0.1 0.000 360.0 360.0 360.0 160.5 -30.2 9.5 -7.7 2 5 A N E -a 105 0A 57 102,-0.9 104,-3.0 122,-0.1 2,-0.5 -0.470 360.0-165.7 -70.5 117.0 -29.5 12.2 -5.1 3 6 A V E +ab 106 124A 3 120,-0.6 122,-2.2 -2,-0.4 123,-1.0 -0.908 13.3 170.6-108.3 127.7 -26.6 14.5 -6.1 4 7 A A E -ab 107 126A 0 102,-2.8 104,-2.6 -2,-0.5 2,-0.5 -0.966 29.8-131.1-136.4 149.1 -25.1 16.8 -3.4 5 8 A I E -ab 108 127A 0 121,-2.7 123,-4.1 -2,-0.3 2,-0.5 -0.881 21.2-167.5 -98.3 130.3 -22.1 19.1 -3.0 6 9 A I E + b 0 128A 3 102,-2.5 2,-0.3 -2,-0.5 123,-0.2 -0.985 22.9 143.1-121.7 125.0 -20.1 18.6 0.2 7 10 A V E - b 0 129A 9 121,-2.6 123,-2.9 -2,-0.5 2,-0.5 -0.989 40.9-137.9-156.3 152.6 -17.5 21.3 1.1 8 11 A A E + b 0 130A 10 -2,-0.3 2,-0.3 121,-0.2 123,-0.2 -0.987 32.4 173.8-118.7 121.6 -15.9 23.2 4.0 9 12 A A E - b 0 131A 0 121,-1.9 123,-2.7 -2,-0.5 2,-0.5 -0.963 32.4-120.0-132.8 145.2 -15.3 26.9 3.5 10 13 A L E >> -Cb 15 132A 8 5,-2.2 4,-1.5 -2,-0.3 5,-1.4 -0.721 51.4 -94.3 -84.0 121.9 -14.1 29.9 5.5 11 14 A K T 45S+ 0 0 69 121,-3.0 123,-0.2 -2,-0.5 -1,-0.0 -0.357 88.2 14.8 -73.7 157.6 -16.7 32.7 5.7 12 15 A P T 45S+ 0 0 112 0, 0.0 -1,-0.2 0, 0.0 121,-0.0 -0.983 133.1 32.1 -91.5 -12.4 -17.5 35.3 4.4 13 16 A A T 45S- 0 0 59 -3,-0.1 133,-0.3 132,-0.0 -2,-0.2 0.582 97.7-129.5 -79.1 -12.3 -15.5 35.0 1.2 14 17 A L T <5 - 0 0 17 -4,-1.5 132,-2.3 1,-0.2 -3,-0.2 0.912 36.2-177.0 60.1 49.7 -15.8 31.1 1.2 15 18 A G E < +CD 10 145A 2 -5,-1.4 -5,-2.2 130,-0.2 130,-0.3 -0.493 16.1 178.2 -73.5 144.9 -12.1 30.4 0.7 16 19 A I E - 0 0 24 128,-2.6 2,-0.3 1,-0.3 129,-0.2 0.441 56.2 -3.6-128.8 -6.3 -11.2 26.7 0.3 17 20 A G E - D 0 144A 4 127,-1.1 126,-2.3 5,-0.2 127,-2.1 -0.985 43.2-143.9-176.3 166.2 -7.4 26.7 -0.2 18 21 A Y B > S-L 21 0B 69 3,-2.5 3,-1.0 -2,-0.3 123,-0.1 -0.976 86.3 -5.1-145.0 132.9 -4.0 28.3 -0.8 19 22 A K T 3 S- 0 0 149 -2,-0.3 3,-0.1 1,-0.2 123,-0.0 0.785 131.4 -53.8 52.9 33.2 -1.2 27.1 -3.0 20 23 A G T 3 S+ 0 0 74 1,-0.2 2,-0.3 0, 0.0 -1,-0.2 0.731 119.4 90.9 78.3 24.8 -3.1 23.9 -3.8 21 24 A K B < S-L 18 0B 138 -3,-1.0 -3,-2.5 0, 0.0 -1,-0.2 -0.985 80.0 -93.5-148.9 159.5 -3.8 22.9 -0.2 22 25 A M - 0 0 119 -2,-0.3 -5,-0.2 -5,-0.2 -6,-0.0 -0.515 23.8-142.9 -76.0 134.2 -6.3 23.2 2.7 23 26 A P S S+ 0 0 32 0, 0.0 2,-0.3 0, 0.0 -1,-0.1 0.195 80.2 39.3 -84.0 17.8 -5.6 26.0 5.2 24 27 A W - 0 0 25 2,-0.0 2,-0.4 155,-0.0 -2,-0.1 -0.987 59.2-148.0-158.5 155.8 -6.8 23.9 8.1 25 28 A R - 0 0 200 -2,-0.3 2,-0.7 4,-0.0 0, 0.0 -0.816 25.0-179.4-128.1 90.6 -6.8 20.4 9.6 26 29 A L > - 0 0 8 -2,-0.4 4,-1.7 1,-0.1 -2,-0.0 -0.834 12.4-161.7 -98.8 111.9 -10.1 20.0 11.5 27 30 A R H > S+ 0 0 142 -2,-0.7 4,-1.1 1,-0.2 -1,-0.1 0.816 87.1 41.6 -67.3 -37.9 -10.3 16.6 13.1 28 31 A K H > S+ 0 0 60 2,-0.2 4,-2.1 1,-0.2 -1,-0.2 0.827 110.0 58.5 -79.6 -33.4 -14.1 16.3 13.6 29 32 A E H > S+ 0 0 18 1,-0.2 4,-2.4 2,-0.2 -2,-0.2 0.884 103.7 53.8 -59.8 -40.6 -14.9 17.8 10.2 30 33 A I H X S+ 0 0 84 -4,-1.7 4,-2.1 2,-0.2 -1,-0.2 0.893 108.7 48.0 -61.1 -44.0 -12.9 15.0 8.6 31 34 A R H X S+ 0 0 123 -4,-1.1 4,-2.7 2,-0.2 -1,-0.2 0.869 110.0 52.8 -64.5 -39.3 -15.0 12.3 10.5 32 35 A Y H X S+ 0 0 7 -4,-2.1 4,-2.5 2,-0.2 5,-0.3 0.925 109.3 48.7 -60.7 -47.8 -18.2 14.1 9.5 33 36 A F H X S+ 0 0 45 -4,-2.4 4,-1.8 1,-0.2 -2,-0.2 0.906 112.9 49.8 -55.5 -44.9 -17.1 14.0 5.8 34 37 A K H X S+ 0 0 88 -4,-2.1 4,-0.6 2,-0.2 -2,-0.2 0.920 114.3 42.1 -62.6 -48.3 -16.3 10.3 6.3 35 38 A D H >X S+ 0 0 75 -4,-2.7 4,-2.7 2,-0.2 3,-0.6 0.925 115.0 47.7 -68.3 -48.1 -19.7 9.3 7.9 36 39 A V H 3< S+ 0 0 2 -4,-2.5 -2,-0.2 1,-0.2 -1,-0.2 0.914 113.3 46.9 -62.9 -46.0 -22.0 11.3 5.6 37 40 A T H 3< S+ 0 0 0 -4,-1.8 -1,-0.2 -5,-0.3 10,-0.2 0.632 119.7 41.8 -71.8 -13.7 -20.4 10.2 2.3 38 41 A T H << S+ 0 0 36 -4,-0.6 9,-0.3 -3,-0.6 -2,-0.2 0.812 86.7 102.6 -99.8 -39.0 -20.4 6.6 3.5 39 42 A R < - 0 0 78 -4,-2.7 2,-0.3 -5,-0.1 7,-0.1 -0.232 47.9-172.7 -59.6 125.4 -23.8 6.1 5.2 40 43 A T - 0 0 45 1,-0.1 -1,-0.0 5,-0.1 -2,-0.0 -0.878 20.4-160.3-115.2 151.5 -26.5 4.3 3.2 41 44 A T S S+ 0 0 81 -2,-0.3 -1,-0.1 2,-0.1 -2,-0.0 0.805 75.2 56.8 -94.2 -38.1 -30.2 3.8 4.0 42 45 A K S > S- 0 0 128 1,-0.1 3,-0.8 4,-0.0 62,-0.1 -0.636 86.9-107.4 -99.5 151.5 -31.0 0.8 1.7 43 46 A P T 3 S+ 0 0 108 0, 0.0 3,-0.1 0, 0.0 -1,-0.1 -0.352 98.0 24.7 -71.9 158.3 -29.3 -2.6 1.6 44 47 A N T 3 S+ 0 0 141 1,-0.2 2,-0.3 -2,-0.1 -2,-0.0 0.585 102.9 105.3 64.4 15.4 -27.0 -3.6 -1.3 45 48 A T < - 0 0 25 -3,-0.8 2,-0.3 58,-0.1 -1,-0.2 -0.800 44.2-175.0-119.8 161.9 -26.1 0.0 -2.1 46 49 A R E -e 103 0A 81 56,-0.8 58,-1.5 -2,-0.3 59,-0.2 -0.960 27.5-104.6-145.9 164.4 -23.2 2.4 -1.6 47 50 A N E - 0 0 0 21,-0.5 23,-2.7 -9,-0.3 2,-0.4 -0.509 25.4-134.0 -88.6 158.0 -22.4 6.0 -2.2 48 51 A A E -ef 106 70A 0 57,-3.1 59,-2.7 21,-0.2 2,-0.5 -0.889 7.9-155.5-109.2 142.4 -20.4 7.6 -4.9 49 52 A V E -ef 107 71A 0 21,-3.5 23,-3.1 -2,-0.4 2,-0.5 -0.981 8.9-163.3-113.3 126.4 -17.8 10.3 -4.6 50 53 A I E +ef 108 72A 1 57,-3.2 59,-2.4 -2,-0.5 60,-0.5 -0.940 19.1 160.8-109.9 132.4 -17.0 12.5 -7.6 51 54 A M E - f 0 73A 0 21,-2.5 23,-2.4 -2,-0.5 60,-0.3 -0.940 38.4-102.2-142.5 161.2 -13.8 14.6 -7.7 52 55 A G E > - f 0 74A 7 58,-1.8 4,-2.0 -2,-0.3 23,-0.2 -0.419 40.0-108.1 -77.0 162.8 -11.5 16.4 -10.1 53 56 A R H > S+ 0 0 97 21,-1.4 4,-2.0 1,-0.2 5,-0.1 0.879 117.8 52.3 -60.0 -43.5 -8.2 14.9 -11.1 54 57 A K H > S+ 0 0 151 1,-0.2 4,-0.7 2,-0.2 -1,-0.2 0.875 108.8 50.6 -62.8 -40.1 -6.1 17.3 -9.1 55 58 A T H >> S+ 0 0 30 2,-0.2 4,-1.0 1,-0.2 3,-0.9 0.912 107.3 54.1 -63.9 -44.5 -8.1 16.6 -5.9 56 59 A W H >< S+ 0 0 36 -4,-2.0 3,-0.9 1,-0.3 -2,-0.2 0.919 109.2 48.2 -53.5 -47.2 -7.6 12.8 -6.5 57 60 A E H 3< S+ 0 0 112 -4,-2.0 -1,-0.3 1,-0.2 -2,-0.2 0.617 106.0 59.0 -72.1 -14.2 -3.8 13.3 -6.7 58 61 A S H << S+ 0 0 79 -3,-0.9 -1,-0.2 -4,-0.7 -2,-0.2 0.664 81.8 101.5 -87.4 -18.6 -3.9 15.4 -3.5 59 62 A I S << S- 0 0 20 -4,-1.0 5,-0.2 -3,-0.9 4,-0.0 -0.478 89.7 -95.5 -69.5 132.1 -5.4 12.6 -1.4 60 63 A P >> - 0 0 77 0, 0.0 3,-2.2 0, 0.0 4,-0.8 -0.179 36.7-113.8 -47.6 138.4 -2.8 10.8 0.8 61 64 A Q G >4 S+ 0 0 136 1,-0.3 3,-0.9 2,-0.2 -2,-0.1 0.839 114.5 43.0 -45.0 -49.7 -1.5 7.6 -0.8 62 65 A K G 34 S+ 0 0 177 1,-0.2 -1,-0.3 4,-0.0 -3,-0.0 0.457 108.6 60.0 -84.9 -0.6 -3.1 5.2 1.7 63 66 A F G <4 S+ 0 0 135 -3,-2.2 -1,-0.2 3,-0.1 -2,-0.2 0.505 86.3 118.2 -98.3 -8.5 -6.4 7.1 1.8 64 67 A R << + 0 0 85 -3,-0.9 2,-0.2 -4,-0.8 -1,-0.0 -0.930 63.1 34.4-116.4 137.0 -7.1 6.6 -1.9 65 68 A P S S- 0 0 47 0, 0.0 22,-0.1 0, 0.0 5,-0.0 0.604 101.1-120.5 -67.3 155.6 -9.3 5.2 -3.2 66 69 A L > - 0 0 19 -2,-0.2 3,-1.0 20,-0.1 -2,-0.1 -0.569 36.6-117.6 -68.7 117.8 -11.6 6.3 -0.3 67 70 A P T 3 S+ 0 0 65 0, 0.0 -1,-0.1 0, 0.0 3,-0.1 -0.117 88.7 20.0 -62.5 153.1 -12.9 3.0 1.2 68 71 A D T 3 S+ 0 0 42 1,-0.1 -21,-0.5 -22,-0.0 2,-0.3 0.524 111.4 87.7 69.8 9.6 -16.6 1.9 1.3 69 72 A R S < S- 0 0 6 -3,-1.0 2,-0.6 -23,-0.1 -21,-0.2 -0.982 80.4-115.3-140.2 146.4 -17.3 4.4 -1.6 70 73 A L E -f 48 0A 10 -23,-2.7 -21,-3.5 -2,-0.3 2,-0.5 -0.738 34.3-145.7 -82.5 122.5 -17.3 4.4 -5.4 71 74 A N E -fg 49 87A 0 15,-3.0 17,-2.5 -2,-0.6 2,-0.5 -0.787 14.7-171.5 -95.8 127.3 -14.7 6.9 -6.6 72 75 A I E -fg 50 88A 1 -23,-3.1 -21,-2.5 -2,-0.5 2,-0.5 -0.978 6.5-162.1-117.7 120.5 -15.3 8.9 -9.8 73 76 A I E -fg 51 89A 0 15,-2.7 17,-2.2 -2,-0.5 2,-0.5 -0.885 9.0-146.0-105.8 132.5 -12.4 10.9 -11.1 74 77 A L E +fg 52 90A 19 -23,-2.4 -21,-1.4 -2,-0.5 2,-0.3 -0.840 28.8 150.6-104.8 129.8 -13.0 13.7 -13.6 75 78 A S > - 0 0 17 15,-1.8 3,-1.2 -2,-0.5 -2,-0.0 -0.968 46.9-133.4-149.2 144.1 -10.7 14.7 -16.4 76 79 A R T 3 S+ 0 0 191 -2,-0.3 15,-0.1 1,-0.2 -2,-0.0 0.485 111.5 55.8 -73.9 -2.1 -11.4 16.3 -19.8 77 80 A S T 3 S+ 0 0 102 13,-0.1 -1,-0.2 2,-0.1 2,-0.1 0.487 87.4 109.0-101.0 -10.7 -9.0 13.6 -21.1 78 81 A Y < - 0 0 53 -3,-1.2 2,-0.3 12,-0.3 13,-0.1 -0.405 59.9-140.4 -79.8 143.7 -10.8 10.6 -19.7 79 82 A E - 0 0 138 -2,-0.1 2,-1.1 11,-0.1 10,-0.1 -0.752 39.3-100.7 -87.0 145.1 -12.8 8.0 -21.5 80 83 A N S S+ 0 0 78 -2,-0.3 2,-0.3 10,-0.1 10,-0.2 -0.581 74.0 140.3 -67.7 99.1 -16.0 6.8 -19.7 81 84 A E E -H 89 0A 83 8,-2.3 8,-2.7 -2,-1.1 2,-0.7 -0.986 57.5-122.1-149.3 152.4 -14.5 3.6 -18.2 82 85 A I E -H 88 0A 108 -2,-0.3 6,-0.2 6,-0.2 3,-0.1 -0.876 22.8-177.0 -99.0 109.0 -14.5 1.4 -15.2 83 86 A I - 0 0 84 4,-1.3 2,-0.2 -2,-0.7 -1,-0.2 0.979 65.5 -35.0 -69.9 -59.7 -10.8 1.0 -14.0 84 87 A D S S- 0 0 73 3,-1.3 -1,-0.2 0, 0.0 3,-0.1 -0.738 81.5 -65.7-143.5-161.8 -11.6 -1.5 -11.1 85 88 A D S S+ 0 0 87 -2,-0.2 3,-0.1 1,-0.2 182,-0.0 0.700 132.1 30.0 -65.8 -21.7 -14.4 -2.2 -8.5 86 89 A N S S+ 0 0 54 1,-0.1 -15,-3.0 -16,-0.1 2,-0.4 0.550 111.4 67.6-114.1 -15.0 -13.8 1.1 -6.7 87 90 A I E +g 71 0A 18 -17,-0.2 -3,-1.3 -3,-0.1 -4,-1.3 -0.942 49.0 178.4-123.4 131.5 -12.6 3.5 -9.4 88 91 A I E -gH 72 82A 7 -17,-2.5 -15,-2.7 -2,-0.4 2,-0.4 -0.986 15.2-150.6-126.4 138.3 -14.3 5.0 -12.4 89 92 A H E +gH 73 81A 32 -8,-2.7 -8,-2.3 -2,-0.4 2,-0.3 -0.924 27.7 161.3-106.8 134.8 -12.9 7.5 -14.9 90 93 A A E -g 74 0A 0 -17,-2.2 -15,-1.8 -2,-0.4 -12,-0.3 -0.972 44.2-143.3-150.0 160.6 -15.2 10.0 -16.6 91 94 A S S S+ 0 0 58 -2,-0.3 2,-0.3 1,-0.2 -17,-0.1 0.370 89.0 13.4-105.4 -0.1 -15.4 13.3 -18.6 92 95 A S > - 0 0 31 1,-0.1 4,-2.1 -17,-0.0 -1,-0.2 -0.969 67.1-119.6-165.2 157.0 -18.6 14.5 -17.0 93 96 A I H > S+ 0 0 5 -2,-0.3 4,-1.8 2,-0.2 5,-0.2 0.829 115.2 61.5 -64.4 -33.4 -21.1 13.8 -14.1 94 97 A E H > S+ 0 0 91 1,-0.2 4,-0.8 2,-0.2 -1,-0.2 0.894 109.9 38.9 -59.8 -43.6 -23.7 13.2 -16.8 95 98 A S H > S+ 0 0 32 2,-0.2 4,-0.8 1,-0.2 3,-0.3 0.894 111.2 56.6 -74.7 -44.1 -21.6 10.2 -18.2 96 99 A S H X S+ 0 0 0 -4,-2.1 4,-1.0 1,-0.2 3,-0.3 0.822 104.6 53.2 -61.1 -35.0 -20.5 8.8 -14.9 97 100 A L H < S+ 0 0 35 -4,-1.8 -1,-0.2 1,-0.2 -2,-0.2 0.859 106.4 52.7 -66.8 -38.0 -24.1 8.4 -13.6 98 101 A N H < S+ 0 0 148 -4,-0.8 -1,-0.2 -3,-0.3 -2,-0.2 0.641 111.6 46.9 -70.8 -16.8 -24.9 6.4 -16.8 99 102 A L H < S+ 0 0 58 -4,-0.8 -1,-0.2 -3,-0.3 -2,-0.2 0.625 96.7 96.0 -95.6 -20.1 -22.0 4.2 -16.0 100 103 A V < - 0 0 27 -4,-1.0 2,-0.4 -5,-0.2 3,-0.1 -0.433 56.6-163.3 -72.5 141.2 -23.0 3.8 -12.4 101 104 A S - 0 0 102 -2,-0.1 -3,-0.0 1,-0.1 -2,-0.0 -0.994 66.7 -8.3-129.2 127.2 -25.1 0.9 -11.1 102 105 A D S S+ 0 0 107 -2,-0.4 -56,-0.8 1,-0.2 2,-0.5 0.890 80.1 156.8 58.7 49.3 -27.0 0.8 -7.8 103 106 A V E - 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