==== Secondary Structure Definition by the program DSSP, updated CMBI version by ElmK / April 1,2000 ==== DATE=31-MAY-2013 . REFERENCE W. KABSCH AND C.SANDER, BIOPOLYMERS 22 (1983) 2577-2637 . HEADER STRUCTURAL PROTEIN 16-OCT-12 4HL8 . COMPND 2 MOLECULE: MAJOR VAULT PROTEIN; . SOURCE 2 ORGANISM_SCIENTIFIC: RATTUS NORVEGICUS; . AUTHOR A.CASANAS,J.QUEROL-AUDI,P.GUERRA,J.POUS,H.TANAKA,T.TSUKIHARA . 782 3 0 0 0 TOTAL NUMBER OF RESIDUES, NUMBER OF CHAINS, NUMBER OF SS-BRIDGES(TOTAL,INTRACHAIN,INTERCHAIN) . 50792.0 ACCESSIBLE SURFACE OF PROTEIN (ANGSTROM**2) . 526 67.3 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(J) , SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS IN PARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 187 23.9 TOTAL NUMBER OF HYDROGEN BONDS IN ANTIPARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 6 0.8 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-5), SAME NUMBER PER 100 RESIDUES . 7 0.9 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-4), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-3), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-2), SAME NUMBER PER 100 RESIDUES . 2 0.3 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-1), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+0), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+1), SAME NUMBER PER 100 RESIDUES . 84 10.7 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+2), SAME NUMBER PER 100 RESIDUES . 41 5.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+3), SAME NUMBER PER 100 RESIDUES . 185 23.7 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+4), SAME NUMBER PER 100 RESIDUES . 4 0.5 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+5), SAME NUMBER PER 100 RESIDUES . 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 *** HISTOGRAMS OF *** . 0 0 0 1 1 1 0 0 0 0 0 0 0 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 2 RESIDUES PER ALPHA HELIX . 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PARALLEL BRIDGES PER LADDER . 16 13 6 4 1 5 3 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 ANTIPARALLEL BRIDGES PER LADDER . 10 4 6 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 LADDERS PER SHEET . # RESIDUE AA STRUCTURE BP1 BP2 ACC N-H-->O O-->H-N N-H-->O O-->H-N TCO KAPPA ALPHA PHI PSI X-CA Y-CA Z-CA 1 5 A E 0 0 196 0, 0.0 32,-0.1 0, 0.0 0, 0.0 0.000 360.0 360.0 360.0 162.6 -86.5 130.1 35.2 2 6 A A + 0 0 39 30,-0.1 31,-1.2 36,-0.1 2,-0.5 0.251 360.0 89.6-100.0 9.1 -88.5 133.4 35.0 3 7 A I B -A 32 0A 107 29,-0.2 2,-0.3 2,-0.0 29,-0.2 -0.938 56.6-172.4-113.2 125.9 -91.2 132.0 37.5 4 8 A I - 0 0 29 27,-2.3 2,-0.6 -2,-0.5 -2,-0.0 -0.862 15.5-145.5-117.2 150.8 -90.7 132.6 41.2 5 9 A R + 0 0 171 -2,-0.3 25,-0.2 25,-0.2 27,-0.0 -0.892 17.8 177.1-120.6 99.0 -92.7 131.1 44.2 6 10 A I B -B 29 0B 5 23,-0.9 23,-0.8 -2,-0.6 25,-0.1 -0.906 20.6-147.2 -98.3 111.5 -93.1 133.5 47.2 7 11 A P > - 0 0 42 0, 0.0 3,-1.0 0, 0.0 22,-0.3 -0.143 39.8 -78.7 -68.0 175.0 -95.3 131.9 49.9 8 12 A P T 3 S+ 0 0 85 0, 0.0 2,-0.1 0, 0.0 19,-0.0 0.744 126.1 27.1 -47.7 -36.5 -97.6 134.0 52.2 9 13 A Y T 3 S+ 0 0 139 17,-0.1 37,-0.7 18,-0.0 2,-0.3 -0.502 99.8 93.4-132.4 64.6 -94.8 135.1 54.5 10 14 A H E < -C 45 0C 54 -3,-1.0 17,-1.3 35,-0.2 2,-0.3 -0.951 46.1-161.6-144.6 164.1 -91.5 135.1 52.6 11 15 A Y E -CD 44 26C 14 33,-2.2 33,-2.0 -2,-0.3 2,-0.3 -0.961 3.9-168.4-145.7 162.6 -89.4 137.7 50.6 12 16 A I E - D 0 25C 0 13,-0.8 13,-1.8 -2,-0.3 2,-0.4 -0.937 22.2-121.2-143.9 164.8 -86.5 137.9 48.1 13 17 A H E - D 0 24C 32 -2,-0.3 27,-2.9 28,-0.3 28,-0.5 -0.946 24.0-164.2-111.9 134.9 -84.2 140.6 46.6 14 18 A V E -ED 39 23C 0 9,-1.4 9,-1.1 -2,-0.4 2,-0.9 -0.977 10.2-153.9-123.9 127.3 -84.2 141.2 42.9 15 19 A L E - D 0 22C 56 23,-2.2 2,-0.7 -2,-0.4 7,-0.2 -0.837 11.9-152.8-101.4 96.0 -81.5 143.1 41.0 16 20 A D E >> - D 0 21C 20 5,-1.0 4,-1.1 -2,-0.9 5,-0.7 -0.601 4.5-159.4 -68.4 112.3 -83.1 144.6 37.8 17 21 A Q T 45S+ 0 0 115 19,-2.3 -1,-0.2 -2,-0.7 20,-0.1 0.735 88.7 48.1 -71.8 -24.2 -80.1 144.8 35.5 18 22 A N T 45S+ 0 0 117 1,-0.1 -1,-0.2 18,-0.1 -2,-0.1 0.906 118.6 38.1 -79.6 -47.0 -81.7 147.5 33.3 19 23 A S T 45S- 0 0 60 2,-0.2 -2,-0.2 56,-0.0 -1,-0.1 0.555 97.3-150.8 -77.7 -10.5 -82.8 149.7 36.2 20 24 A N T <5 + 0 0 124 -4,-1.1 2,-0.3 1,-0.1 -3,-0.2 0.724 64.4 87.7 45.1 35.1 -79.5 148.8 37.9 21 25 A V E < -D 16 0C 19 -5,-0.7 -5,-1.0 54,-0.1 2,-0.2 -0.865 69.3-132.5-145.5 168.0 -80.9 149.2 41.4 22 26 A S E -D 15 0C 14 -2,-0.3 2,-0.3 -7,-0.2 55,-0.2 -0.719 16.5-169.1-121.2 173.1 -82.7 146.6 43.5 23 27 A R E -D 14 0C 57 -9,-1.1 -9,-1.4 -2,-0.2 2,-0.2 -0.976 30.8 -90.5-156.0 165.2 -85.9 146.4 45.6 24 28 A V E -D 13 0C 25 -2,-0.3 2,-0.4 -11,-0.2 -11,-0.2 -0.557 26.9-154.7 -80.5 145.1 -87.7 144.2 48.1 25 29 A E E +D 12 0C 28 -13,-1.8 -13,-0.8 -2,-0.2 2,-0.2 -0.719 19.2 178.4-123.7 79.0 -90.2 141.6 46.9 26 30 A V E -D 11 0C 67 -2,-0.4 -15,-0.2 -15,-0.2 -17,-0.1 -0.587 19.3-143.0 -83.3 142.2 -92.7 140.9 49.6 27 31 A G S S+ 0 0 15 -17,-1.3 -1,-0.1 -2,-0.2 3,-0.0 0.116 71.4 136.4 -78.4-162.4 -95.5 138.4 49.1 28 32 A P S S+ 0 0 102 0, 0.0 2,-0.3 0, 0.0 -1,-0.1 0.202 83.5 86.8 -86.1 112.0 -98.1 136.9 49.0 29 33 A K B S-B 6 0B 114 -23,-0.8 2,-1.7 -22,-0.3 -23,-0.9 -0.896 71.9-128.7-167.6 140.5 -97.7 136.2 45.3 30 34 A T - 0 0 67 -2,-0.3 -25,-0.2 -25,-0.2 -26,-0.1 -0.576 51.4-165.8 -83.4 72.7 -96.0 133.6 42.9 31 35 A Y - 0 0 67 -2,-1.7 -27,-2.3 -28,-0.1 2,-0.5 -0.198 15.3-140.0 -69.9 151.8 -94.4 136.5 41.1 32 36 A I B -A 3 0A 102 -29,-0.2 -29,-0.2 -27,-0.0 2,-0.1 -0.959 16.7-133.8-115.1 124.3 -92.8 136.3 37.6 33 37 A R - 0 0 68 -31,-1.2 2,-0.1 -2,-0.5 -19,-0.0 -0.417 18.5-123.2 -75.0 149.6 -89.5 138.1 37.0 34 38 A Q - 0 0 82 -2,-0.1 -1,-0.1 1,-0.1 0, 0.0 -0.264 32.4 -98.5 -79.0 174.5 -89.0 140.2 33.8 35 39 A D S S+ 0 0 153 1,-0.2 -1,-0.1 -2,-0.1 -2,-0.1 0.796 127.6 47.4 -63.7 -29.7 -86.1 139.6 31.3 36 40 A N S S+ 0 0 62 -20,-0.1 -19,-2.3 -19,-0.1 -1,-0.2 0.833 100.7 82.3 -79.5 -36.3 -84.2 142.5 33.0 37 41 A E - 0 0 11 -21,-0.3 2,-0.3 -20,-0.1 -21,-0.2 -0.234 54.2-175.7 -69.7 157.3 -84.9 141.2 36.5 38 42 A R - 0 0 112 -36,-0.0 -23,-2.2 0, 0.0 2,-0.2 -0.845 22.4-131.4-156.9 118.7 -82.8 138.5 38.3 39 43 A V B -E 14 0C 32 -2,-0.3 -25,-0.2 -25,-0.2 3,-0.1 -0.522 11.6-159.7 -75.3 133.9 -83.6 137.1 41.7 40 44 A L + 0 0 88 -27,-2.9 2,-0.3 -2,-0.2 -1,-0.1 0.932 67.1 0.4 -79.0 -49.6 -80.6 136.9 44.1 41 45 A F S S- 0 0 70 -28,-0.5 -28,-0.3 1,-0.1 -1,-0.2 -0.924 80.3 -87.2-143.7 163.9 -81.7 134.3 46.6 42 46 A A - 0 0 68 -2,-0.3 -31,-0.1 1,-0.2 -29,-0.1 -0.127 65.0 -67.2 -69.7 164.1 -84.6 132.0 47.4 43 47 A P - 0 0 47 0, 0.0 2,-0.4 0, 0.0 -31,-0.2 -0.316 58.1-173.8 -51.5 126.7 -87.7 133.0 49.5 44 48 A V E -C 11 0C 67 -33,-2.0 -33,-2.2 -3,-0.1 -3,-0.0 -0.958 33.2 -93.6-123.7 145.1 -86.4 133.5 53.1 45 49 A R E -C 10 0C 126 -2,-0.4 -35,-0.2 -35,-0.2 3,-0.1 -0.252 45.1-102.9 -60.7 142.9 -88.8 134.1 56.0 46 50 A M - 0 0 18 -37,-0.7 2,-0.6 1,-0.1 -1,-0.1 -0.332 48.8 -85.7 -66.8 147.7 -89.5 137.8 56.9 47 51 A V E -F 93 0D 16 46,-2.0 46,-2.0 45,-0.1 2,-0.2 -0.396 54.0-164.4 -63.4 105.3 -87.8 139.2 60.0 48 52 A T E - 0 0 32 -2,-0.6 57,-0.3 44,-0.3 43,-0.1 -0.551 14.6-173.8 -89.2 155.0 -90.0 138.2 62.9 49 53 A V E -F 90 0D 6 41,-0.9 41,-1.3 -2,-0.2 3,-0.1 -0.741 15.8-147.9-150.8 97.9 -89.9 139.7 66.4 50 54 A P > - 0 0 47 0, 0.0 3,-1.3 0, 0.0 2,-0.1 -0.037 38.2 -75.4 -58.5 165.6 -92.1 138.1 69.1 51 55 A P T 3 S+ 0 0 103 0, 0.0 3,-0.1 0, 0.0 38,-0.1 -0.380 118.6 16.8 -63.5 143.9 -93.6 140.1 72.0 52 56 A R T 3 S+ 0 0 199 1,-0.3 56,-1.3 -3,-0.1 2,-0.2 0.667 113.2 92.6 63.1 21.4 -91.2 141.0 74.8 53 57 A H E < -G 107 0E 42 -3,-1.3 2,-0.3 54,-0.2 -1,-0.3 -0.586 58.4-154.9-122.0-173.4 -88.2 140.2 72.5 54 58 A Y E -G 106 0E 21 52,-1.6 52,-1.7 -2,-0.2 2,-0.3 -0.991 12.7-167.4-159.5 163.6 -86.2 142.5 70.1 55 59 A C B -H 87 0F 0 32,-2.4 32,-1.3 -2,-0.3 2,-0.3 -0.969 7.2-158.5-149.2 161.5 -84.0 142.7 67.0 56 60 A I - 0 0 11 47,-0.4 47,-2.0 -2,-0.3 46,-0.9 -0.945 5.5-168.5-148.6 117.1 -81.7 145.4 65.6 57 61 A V B -I 101 0G 0 28,-0.4 28,-0.8 -2,-0.3 44,-0.2 -0.938 15.1-139.8-114.4 126.1 -80.6 145.7 61.9 58 62 A A B S+J 84 0H 11 42,-1.5 26,-0.2 -2,-0.5 42,-0.1 -0.213 82.9 41.6 -74.2 171.5 -77.8 148.0 60.8 59 63 A N S S- 0 0 42 24,-1.5 -1,-0.2 22,-1.2 25,-0.2 0.903 87.0-164.4 55.0 46.7 -78.0 150.0 57.5 60 64 A P - 0 0 0 0, 0.0 20,-2.5 0, 0.0 2,-0.6 -0.294 26.3-104.1 -70.0 143.7 -81.7 150.8 58.2 61 65 A V B -K 79 0I 0 18,-0.2 2,-0.6 19,-0.1 18,-0.3 -0.535 39.3-127.7 -65.0 109.9 -84.0 152.1 55.5 62 66 A S - 0 0 23 16,-1.8 8,-0.6 -2,-0.6 2,-0.3 -0.496 30.2-166.4 -67.2 109.5 -84.4 155.8 56.4 63 67 A R B -L 69 0J 74 -2,-0.6 6,-0.2 6,-0.1 -1,-0.0 -0.725 15.5-122.8 -98.2 147.8 -88.1 156.6 56.6 64 68 A D > - 0 0 69 4,-0.7 3,-1.8 -2,-0.3 -1,-0.1 0.042 49.6 -74.9 -71.1-171.8 -89.6 160.1 56.7 65 69 A T T 3 S+ 0 0 152 1,-0.3 -1,-0.1 2,-0.1 -2,-0.0 0.609 133.6 57.4 -65.4 -11.9 -91.9 161.4 59.5 66 70 A Q T 3 S- 0 0 138 2,-0.1 -1,-0.3 0, 0.0 -2,-0.0 0.463 113.1-117.5 -95.0 -4.3 -94.7 159.3 58.0 67 71 A S < + 0 0 89 -3,-1.8 2,-0.3 1,-0.2 -2,-0.1 0.730 60.6 148.6 71.5 26.7 -92.7 156.0 58.3 68 72 A S - 0 0 58 1,-0.0 2,-1.4 -6,-0.0 -4,-0.7 -0.715 57.5-109.9 -93.8 143.4 -92.5 155.1 54.6 69 73 A V B -L 63 0J 35 -2,-0.3 2,-0.4 -6,-0.2 -6,-0.1 -0.534 35.6-146.0 -73.0 93.1 -89.7 153.3 52.9 70 74 A L - 0 0 46 -2,-1.4 8,-2.3 -8,-0.6 2,-0.5 -0.482 8.0-150.7 -63.3 115.4 -88.2 156.1 50.8 71 75 A F B -M 77 0K 99 -2,-0.4 6,-0.3 6,-0.2 5,-0.1 -0.825 25.8-114.8 -87.1 129.1 -87.0 154.5 47.6 72 76 A D > - 0 0 63 4,-3.4 3,-0.7 -2,-0.5 -1,-0.1 -0.119 23.6-104.9 -63.8 162.2 -84.1 156.6 46.4 73 77 A I T 3 S+ 0 0 172 1,-0.3 -1,-0.1 2,-0.1 4,-0.1 0.817 124.7 55.4 -60.7 -32.3 -84.3 158.5 43.1 74 78 A T T 3 S- 0 0 88 2,-0.1 -1,-0.3 1,-0.0 3,-0.1 0.852 123.4-108.8 -63.4 -36.0 -82.1 155.8 41.5 75 79 A G S < S+ 0 0 24 -3,-0.7 2,-0.1 1,-0.5 -2,-0.1 0.173 82.1 118.1 122.4 -13.4 -84.6 153.2 42.7 76 80 A Q S S- 0 0 48 -5,-0.1 -4,-3.4 1,-0.1 -1,-0.5 -0.458 77.4 -91.8 -76.1 154.4 -82.6 151.6 45.4 77 81 A V B -M 71 0K 5 -6,-0.3 2,-0.8 -55,-0.2 -6,-0.2 -0.483 41.3-115.1 -67.8 137.0 -83.9 151.8 49.0 78 82 A R - 0 0 106 -8,-2.3 -16,-1.8 -2,-0.2 2,-0.3 -0.671 40.9-172.6 -80.2 110.3 -82.5 154.8 50.9 79 83 A L B -K 61 0I 22 -2,-0.8 -18,-0.2 -18,-0.3 3,-0.1 -0.753 32.7-144.1-106.6 148.6 -80.4 153.4 53.7 80 84 A R > - 0 0 90 -20,-2.5 3,-0.5 1,-0.3 2,-0.4 0.619 31.8-167.8 -80.6 -15.7 -78.7 155.3 56.7 81 85 A H T 3 S+ 0 0 117 1,-0.3 -22,-1.2 -21,-0.2 -1,-0.3 -0.515 75.3 39.3 64.1-120.5 -75.7 152.9 56.3 82 86 A A T 3 S+ 0 0 103 -2,-0.4 -1,-0.3 -23,-0.1 -2,-0.0 0.590 105.0 77.3 -37.1 -32.8 -73.9 153.7 59.6 83 87 A D < - 0 0 61 -3,-0.5 -24,-1.5 -23,-0.1 2,-0.3 -0.042 67.2-144.4 -73.5 176.5 -77.0 154.0 61.8 84 88 A Q B -J 58 0H 53 -26,-0.2 2,-0.4 -25,-0.2 -26,-0.2 -0.860 6.8-127.7-133.9 171.1 -79.1 151.2 63.3 85 89 A E - 0 0 35 -28,-0.8 2,-1.0 -2,-0.3 -28,-0.4 -0.961 11.2-136.2-124.4 146.4 -82.8 150.7 64.1 86 90 A I - 0 0 72 -2,-0.4 2,-0.8 -30,-0.2 -30,-0.2 -0.721 22.5-170.7-103.2 83.6 -84.4 149.5 67.4 87 91 A R B +H 55 0F 13 -32,-1.3 -32,-2.4 -2,-1.0 3,-0.1 -0.647 6.4 179.1 -78.7 110.7 -87.0 146.9 66.3 88 92 A L - 0 0 104 -2,-0.8 2,-0.3 1,-0.3 -35,-0.2 0.990 57.7 -42.9 -76.6 -73.5 -89.1 146.2 69.4 89 93 A A S S+ 0 0 46 -35,-0.1 2,-0.3 -37,-0.1 -1,-0.3 -0.907 78.9 132.7-163.2 133.3 -91.8 143.6 68.6 90 94 A Q E -F 49 0D 101 -41,-1.3 -41,-0.9 -2,-0.3 3,-0.1 -0.941 54.6 -57.5-172.4 179.1 -93.8 143.8 65.4 91 95 A D E S- 0 0 127 -2,-0.3 2,-0.8 1,-0.2 -44,-0.0 -0.131 72.4 -72.6 -68.9 166.0 -95.1 141.6 62.5 92 96 A P E - 0 0 36 0, 0.0 -44,-0.3 0, 0.0 -1,-0.2 -0.512 62.3-173.1 -60.2 104.4 -92.8 139.6 60.1 93 97 A F E -F 47 0D 24 -46,-2.0 -46,-2.0 -2,-0.8 2,-0.3 -0.819 20.8-119.3-104.5 144.6 -91.3 142.4 58.0 94 98 A P - 0 0 15 0, 0.0 2,-0.1 0, 0.0 -48,-0.1 -0.616 23.1-124.5 -84.0 139.4 -89.1 141.9 55.0 95 99 A L - 0 0 12 -2,-0.3 3,-0.1 1,-0.1 -83,-0.1 -0.434 24.3-129.5 -74.0 156.4 -85.5 143.3 54.9 96 100 A Y > - 0 0 24 1,-0.1 3,-1.2 -2,-0.1 -83,-0.1 -0.539 38.7 -78.0-103.4 169.1 -84.5 145.5 52.0 97 101 A P T 3 S+ 0 0 10 0, 0.0 -1,-0.1 0, 0.0 -20,-0.1 -0.455 120.6 31.8 -66.5 147.2 -81.5 145.2 49.7 98 102 A G T 3 S+ 0 0 45 1,-0.1 2,-0.6 -3,-0.1 -17,-0.1 0.613 82.5 121.7 81.2 6.3 -78.6 146.5 51.9 99 103 A E < - 0 0 15 -3,-1.2 2,-0.7 -20,-0.0 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