==== Secondary Structure Definition by the program DSSP, updated CMBI version by ElmK / April 1,2000 ==== DATE=4-DEC-2009 . REFERENCE W. KABSCH AND C.SANDER, BIOPOLYMERS 22 (1983) 2577-2637 . HEADER TOXIN 28-FEB-01 1I5P . COMPND 2 MOLECULE: PESTICIDIAL CRYSTAL PROTEIN CRY2AA; . SOURCE 2 ORGANISM_SCIENTIFIC: BACILLUS THURINGIENSIS SEROVAR . AUTHOR R.J.MORSE,R.M.STROUD,T.YAMAMOTO . 633 1 1 1 0 TOTAL NUMBER OF RESIDUES, NUMBER OF CHAINS, NUMBER OF SS-BRIDGES(TOTAL,INTRACHAIN,INTERCHAIN) . 22995.0 ACCESSIBLE SURFACE OF PROTEIN (ANGSTROM**2) . 446 70.5 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(J) , SAME NUMBER PER 100 RESIDUES . 12 1.9 TOTAL NUMBER OF HYDROGEN BONDS IN PARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 133 21.0 TOTAL NUMBER OF HYDROGEN BONDS IN ANTIPARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-5), SAME NUMBER PER 100 RESIDUES . 4 0.6 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-4), SAME NUMBER PER 100 RESIDUES . 2 0.3 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-3), SAME NUMBER PER 100 RESIDUES . 1 0.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-2), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-1), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+0), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+1), SAME NUMBER PER 100 RESIDUES . 65 10.3 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+2), SAME NUMBER PER 100 RESIDUES . 46 7.3 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+3), SAME NUMBER PER 100 RESIDUES . 164 25.9 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+4), SAME NUMBER PER 100 RESIDUES . 5 0.8 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+5), SAME NUMBER PER 100 RESIDUES . 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 *** HISTOGRAMS OF *** . 0 0 0 3 0 0 1 2 0 0 0 0 1 3 0 1 0 0 0 0 0 0 0 1 0 0 0 0 1 1 RESIDUES PER ALPHA HELIX . 1 2 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PARALLEL BRIDGES PER LADDER . 2 4 6 3 2 1 0 3 1 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 ANTIPARALLEL BRIDGES PER LADDER . 2 1 1 0 0 2 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 LADDERS PER SHEET . # RESIDUE AA STRUCTURE BP1 BP2 ACC N-H-->O O-->H-N N-H-->O O-->H-N TCO KAPPA ALPHA PHI PSI X-CA Y-CA Z-CA 1 1 A M 0 0 158 0, 0.0 4,-0.1 0, 0.0 2,-0.0 0.000 360.0 360.0 360.0 179.1 -33.0 -64.7 117.1 2 2 A N - 0 0 79 474,-0.1 475,-0.4 472,-0.0 367,-0.1 0.177 360.0 -10.0-117.5-124.8 -30.4 -62.2 116.1 3 3 A N S S+ 0 0 21 1,-0.3 2,-1.7 473,-0.1 366,-0.2 0.717 116.7 83.8 -53.7 -20.4 -26.9 -62.6 114.7 4 4 A V + 0 0 27 365,-0.1 2,-0.4 366,-0.1 -1,-0.3 -0.155 67.7 169.0 -77.4 44.0 -27.3 -66.2 115.6 5 5 A L - 0 0 21 -2,-1.7 364,-1.4 362,-0.2 365,-0.5 -0.506 30.8-122.5 -65.2 117.7 -29.1 -66.6 112.3 6 6 A N + 0 0 91 -2,-0.4 2,-0.3 361,-0.3 360,-0.1 -0.261 50.4 122.4 -76.3 147.9 -29.5 -70.4 111.7 7 7 A S + 0 0 27 358,-0.3 2,-0.2 357,-0.2 356,-0.0 -0.893 24.3 165.6-179.8 168.3 -28.4 -72.9 109.0 8 8 A G - 0 0 43 -2,-0.3 357,-0.3 1,-0.1 356,-0.2 -0.817 35.2-152.9 167.7 152.3 -26.3 -76.1 109.0 9 9 A R - 0 0 38 354,-0.4 355,-0.1 -2,-0.2 -1,-0.1 0.816 64.4 -85.8-105.4 -47.6 -25.1 -79.3 107.4 10 10 A T S S+ 0 0 107 1,-0.3 352,-0.1 353,-0.2 354,-0.0 0.047 76.1 124.2 171.1 -50.1 -24.1 -81.7 110.3 11 11 A T - 0 0 58 1,-0.1 -1,-0.3 394,-0.1 2,-0.1 -0.082 52.6-135.4 -44.3 125.7 -20.6 -81.5 111.9 12 12 A I + 0 0 154 1,-0.1 -1,-0.1 353,-0.1 353,-0.0 -0.465 46.7 140.7 -86.3 155.7 -20.9 -80.9 115.6 13 13 A C - 0 0 47 -2,-0.1 -1,-0.1 391,-0.0 -2,-0.1 0.258 55.9-119.7-172.7 3.5 -18.8 -78.4 117.7 14 14 A D - 0 0 118 1,-0.1 4,-0.2 3,-0.1 5,-0.1 0.621 16.2-106.2 54.9 145.9 -21.3 -77.0 120.3 15 15 A A S >> S+ 0 0 39 1,-0.2 2,-3.3 2,-0.2 4,-2.7 0.840 106.6 82.6 -68.1 -35.8 -22.4 -73.4 120.7 16 16 A Y T 34 S+ 0 0 215 1,-0.3 -1,-0.2 2,-0.2 0, 0.0 -0.375 99.1 37.9 -70.8 68.6 -20.4 -73.2 123.9 17 17 A N T 34 S+ 0 0 115 -2,-3.3 -1,-0.3 3,-0.0 -2,-0.2 0.157 137.3 11.1 173.8 -20.9 -17.2 -72.5 121.9 18 18 A V T <4 S+ 0 0 27 -3,-2.1 -2,-0.2 -4,-0.2 -3,-0.1 0.130 82.4 121.9-160.1 19.6 -18.5 -70.3 119.0 19 19 A V < + 0 0 80 -4,-2.7 2,-0.4 -5,-0.1 -3,-0.1 0.815 70.0 60.7 -61.2 -31.6 -22.0 -69.3 120.0 20 20 A A - 0 0 87 -5,-0.3 2,-0.4 -16,-0.0 -1,-0.0 -0.845 60.8-177.2-106.7 135.7 -21.2 -65.6 119.8 21 21 A H - 0 0 31 -2,-0.4 -18,-0.1 350,-0.0 350,-0.1 -0.993 23.9-133.0-128.1 133.2 -20.0 -63.6 116.8 22 22 A D > - 0 0 59 -2,-0.4 3,-1.7 1,-0.1 4,-0.2 -0.774 10.2-164.7 -92.0 108.8 -19.0 -60.0 116.8 23 23 A P T 3 S+ 0 0 21 0, 0.0 449,-0.2 0, 0.0 -1,-0.1 0.535 94.5 43.9 -66.9 -6.1 -20.6 -58.1 113.9 24 24 A F T 3> S+ 0 0 6 1,-0.1 4,-2.2 2,-0.1 5,-0.2 -0.108 72.0 131.0-127.6 32.6 -18.2 -55.3 114.5 25 25 A S H <> S+ 0 0 28 -3,-1.7 4,-0.7 1,-0.2 -1,-0.1 0.897 74.7 49.9 -54.7 -44.2 -15.0 -57.3 115.0 26 26 A F H >4 S+ 0 0 23 1,-0.2 3,-1.1 2,-0.2 -1,-0.2 0.949 112.2 47.0 -59.8 -49.8 -13.0 -55.2 112.6 27 27 A E H 34 S+ 0 0 65 1,-0.2 -1,-0.2 -4,-0.1 -2,-0.2 0.827 112.8 50.5 -62.1 -30.6 -14.1 -52.0 114.3 28 28 A H H 3< S+ 0 0 112 -4,-2.2 2,-0.3 -6,-0.1 -1,-0.2 0.535 93.8 95.9 -85.4 -6.9 -13.3 -53.5 117.8 29 29 A K S << S- 0 0 46 -3,-1.1 2,-0.1 -4,-0.7 -3,-0.0 -0.619 73.9-127.1 -86.9 142.5 -9.8 -54.6 116.7 30 30 A S > - 0 0 59 -2,-0.3 4,-2.1 1,-0.1 5,-0.2 -0.456 23.2-115.9 -81.1 160.8 -6.8 -52.5 117.4 31 31 A L H > S+ 0 0 48 1,-0.2 4,-3.2 2,-0.2 5,-0.2 0.887 116.0 57.0 -65.6 -37.2 -4.5 -51.5 114.5 32 32 A D H > S+ 0 0 87 1,-0.2 4,-2.2 2,-0.2 -1,-0.2 0.907 107.7 47.4 -59.6 -42.5 -1.6 -53.5 116.1 33 33 A T H > S+ 0 0 54 2,-0.2 4,-2.3 1,-0.2 -1,-0.2 0.926 113.8 47.7 -64.1 -44.4 -3.8 -56.6 116.1 34 34 A I H X S+ 0 0 0 -4,-2.1 4,-2.3 2,-0.2 -2,-0.2 0.912 110.4 51.6 -63.1 -43.2 -4.8 -56.0 112.4 35 35 A Q H X S+ 0 0 62 -4,-3.2 4,-2.2 1,-0.2 -1,-0.2 0.915 110.8 49.1 -60.8 -42.2 -1.2 -55.4 111.4 36 36 A K H X S+ 0 0 118 -4,-2.2 4,-2.6 -5,-0.2 -1,-0.2 0.883 109.1 52.4 -64.5 -39.9 -0.2 -58.7 113.1 37 37 A E H X S+ 0 0 58 -4,-2.3 4,-2.5 2,-0.2 -1,-0.2 0.909 111.1 46.0 -64.2 -41.3 -3.0 -60.6 111.3 38 38 A W H X S+ 0 0 0 -4,-2.3 4,-2.5 2,-0.2 -1,-0.2 0.889 112.6 51.2 -68.3 -37.8 -1.9 -59.3 107.9 39 39 A M H X S+ 0 0 122 -4,-2.2 4,-1.9 2,-0.2 -2,-0.2 0.913 113.0 45.0 -64.2 -42.9 1.7 -60.1 108.7 40 40 A E H X S+ 0 0 90 -4,-2.6 4,-2.6 2,-0.2 5,-0.4 0.892 109.8 55.6 -67.5 -38.8 0.7 -63.6 109.7 41 41 A W H X S+ 0 0 2 -4,-2.5 4,-2.9 1,-0.2 -2,-0.2 0.942 107.7 50.0 -56.4 -47.5 -1.4 -63.9 106.7 42 42 A K H < S+ 0 0 48 -4,-2.5 5,-0.3 1,-0.2 -1,-0.2 0.919 113.1 46.0 -57.6 -46.2 1.6 -63.1 104.6 43 43 A R H < S+ 0 0 174 -4,-1.9 -2,-0.2 1,-0.1 -1,-0.2 0.879 129.6 17.7 -65.8 -43.5 3.7 -65.7 106.3 44 44 A T H < S+ 0 0 77 -4,-2.6 -3,-0.2 -5,-0.1 -2,-0.2 0.642 112.2 67.4-108.1 -19.9 1.3 -68.6 106.2 45 45 A D S < S- 0 0 31 -4,-2.9 2,-1.5 -5,-0.4 371,-0.1 -0.470 90.7 -85.3-105.6 175.9 -1.4 -67.9 103.6 46 46 A H + 0 0 35 369,-0.6 371,-0.3 -2,-0.1 2,-0.2 -0.632 51.0 175.5 -77.8 92.2 -1.8 -67.5 99.9 47 47 A S - 0 0 21 -2,-1.5 4,-0.1 -5,-0.3 371,-0.0 -0.585 37.6-127.8 -88.1 161.2 -0.9 -63.9 99.5 48 48 A L - 0 0 4 -2,-0.2 3,-0.1 198,-0.1 -2,-0.1 -0.108 67.4 -76.6-102.1 39.1 -0.9 -62.9 95.8 49 49 A Y S S+ 0 0 103 1,-0.3 2,-0.3 -7,-0.1 51,-0.2 0.637 93.6 155.8 73.6 16.1 2.6 -61.5 96.4 50 50 A V - 0 0 0 1,-0.1 -1,-0.3 50,-0.0 43,-0.1 -0.603 48.9-121.4 -76.7 131.2 0.6 -58.7 98.0 51 51 A A > - 0 0 2 -2,-0.3 3,-2.1 1,-0.1 4,-0.5 -0.647 15.8-137.5 -75.5 121.4 2.5 -56.8 100.6 52 52 A P T >> S+ 0 0 24 0, 0.0 4,-1.5 0, 0.0 3,-1.0 0.755 96.5 73.9 -50.7 -30.2 0.7 -57.1 103.9 53 53 A V H 3> S+ 0 0 8 1,-0.3 4,-3.1 2,-0.2 5,-0.2 0.818 86.0 64.3 -56.6 -29.9 1.2 -53.4 104.6 54 54 A V H <> S+ 0 0 0 -3,-2.1 4,-2.3 1,-0.2 -1,-0.3 0.897 102.7 48.8 -62.0 -36.3 -1.4 -52.6 102.0 55 55 A G H <> S+ 0 0 0 -3,-1.0 4,-2.2 -4,-0.5 -1,-0.2 0.894 111.7 48.4 -68.8 -40.3 -4.0 -54.3 104.2 56 56 A T H X S+ 0 0 2 -4,-1.5 4,-2.7 2,-0.2 -2,-0.2 0.937 112.4 48.3 -64.9 -47.2 -2.8 -52.4 107.3 57 57 A V H X S+ 0 0 0 -4,-3.1 4,-2.9 2,-0.2 5,-0.2 0.925 112.9 46.8 -60.8 -46.6 -2.9 -49.1 105.6 58 58 A S H X S+ 0 0 4 -4,-2.3 4,-2.7 -5,-0.2 -1,-0.2 0.931 113.4 48.7 -62.6 -43.7 -6.4 -49.6 104.1 59 59 A S H X S+ 0 0 0 -4,-2.2 4,-2.1 1,-0.2 -1,-0.2 0.917 112.0 51.4 -60.0 -42.2 -7.7 -50.8 107.5 60 60 A F H X S+ 0 0 6 -4,-2.7 4,-1.7 2,-0.2 -2,-0.2 0.922 110.7 46.0 -60.1 -48.9 -6.0 -47.8 109.1 61 61 A L H X S+ 0 0 1 -4,-2.9 4,-2.3 1,-0.2 5,-0.3 0.914 109.4 55.3 -63.9 -41.4 -7.7 -45.4 106.6 62 62 A L H X S+ 0 0 13 -4,-2.7 4,-1.9 1,-0.2 207,-0.3 0.911 110.1 47.2 -56.5 -42.4 -11.1 -47.1 107.1 63 63 A K H < S+ 0 0 54 -4,-2.1 4,-0.3 2,-0.2 -1,-0.2 0.799 108.7 53.7 -70.7 -30.7 -10.8 -46.5 110.8 64 64 A K H >< S+ 0 0 23 -4,-1.7 3,-0.8 1,-0.2 6,-0.5 0.926 115.6 39.3 -68.2 -45.0 -9.7 -42.8 110.4 65 65 A V H 3< S+ 0 0 0 -4,-2.3 202,-3.0 1,-0.2 201,-0.4 0.808 110.9 61.4 -72.5 -31.4 -12.8 -42.1 108.3 66 66 A G T 3< S+ 0 0 16 -4,-1.9 2,-0.4 -5,-0.3 -1,-0.2 0.496 105.6 50.0 -76.1 -3.1 -14.9 -44.3 110.5 67 67 A S S X S- 0 0 45 -3,-0.8 3,-2.4 -4,-0.3 199,-0.2 -0.991 70.7-138.9-142.8 133.5 -14.3 -42.2 113.6 68 68 A L T 3 S+ 0 0 70 -2,-0.4 3,-0.1 1,-0.3 -1,-0.1 0.746 102.9 70.0 -57.8 -22.8 -14.6 -38.5 114.3 69 69 A I T 3 S+ 0 0 143 1,-0.2 2,-2.0 2,-0.1 3,-0.4 0.702 74.3 89.4 -69.8 -17.5 -11.4 -38.8 116.3 70 70 A G < + 0 0 1 -3,-2.4 -1,-0.2 -6,-0.5 3,-0.1 -0.263 44.1 134.2 -79.3 54.9 -9.5 -39.4 113.1 71 71 A K + 0 0 73 -2,-2.0 2,-0.5 1,-0.2 -1,-0.2 0.778 67.5 44.5 -72.3 -30.0 -8.8 -35.7 112.5 72 72 A R + 0 0 53 -3,-0.4 4,-0.2 1,-0.1 -1,-0.2 -0.966 55.7 157.5-125.3 121.0 -5.1 -36.2 111.6 73 73 A I > + 0 0 0 -2,-0.5 4,-2.8 113,-0.4 3,-0.4 0.838 59.1 71.1-104.4 -48.3 -4.0 -39.0 109.3 74 74 A L H > S+ 0 0 0 112,-0.4 4,-2.4 1,-0.2 5,-0.2 0.782 95.4 52.5 -45.0 -44.8 -0.6 -38.1 107.8 75 75 A S H > S+ 0 0 42 2,-0.2 4,-1.3 1,-0.2 -1,-0.2 0.945 117.4 36.1 -62.9 -48.8 1.6 -38.6 111.0 76 76 A E H > S+ 0 0 109 -3,-0.4 4,-0.8 -4,-0.2 -2,-0.2 0.875 114.7 57.3 -72.6 -35.3 0.4 -42.1 111.8 77 77 A L H >X S+ 0 0 0 -4,-2.8 4,-2.5 1,-0.2 3,-1.5 0.943 104.6 52.1 -57.9 -47.8 0.1 -43.0 108.2 78 78 A W H 3X S+ 0 0 45 -4,-2.4 4,-2.1 1,-0.3 7,-0.3 0.875 105.3 56.6 -54.7 -40.4 3.8 -42.2 107.7 79 79 A G H 3< S+ 0 0 38 -4,-1.3 -1,-0.3 -5,-0.2 5,-0.2 0.670 112.6 40.8 -68.1 -18.0 4.6 -44.4 110.7 80 80 A I H << S+ 0 0 12 -3,-1.5 -2,-0.2 -4,-0.8 -1,-0.2 0.875 121.6 36.1 -94.1 -50.9 3.0 -47.4 109.0 81 81 A I H < S+ 0 0 0 -4,-2.5 9,-0.3 1,-0.3 8,-0.2 0.721 130.5 34.3 -77.3 -22.7 4.1 -47.1 105.4 82 82 A F S >< S- 0 0 0 -4,-2.1 3,-2.8 -5,-0.4 -1,-0.3 -0.600 87.2-156.1-129.4 66.9 7.5 -45.8 106.4 83 83 A P G > S- 0 0 42 0, 0.0 3,-1.7 0, 0.0 5,-0.2 -0.280 79.0 -18.3 -52.0 110.5 8.3 -47.7 109.6 84 84 A S G 3 S- 0 0 128 1,-0.3 -5,-0.1 -2,-0.2 -4,-0.0 0.767 129.0 -55.2 59.7 29.6 10.8 -45.6 111.5 85 85 A G G < S+ 0 0 20 -3,-2.8 -1,-0.3 -7,-0.3 2,-0.2 0.457 100.8 143.1 87.0 1.2 11.6 -43.8 108.3 86 86 A S < - 0 0 46 -3,-1.7 3,-0.3 1,-0.1 -1,-0.3 -0.534 41.8-159.5 -80.2 141.6 12.4 -46.9 106.3 87 87 A T > + 0 0 20 -2,-0.2 4,-2.6 1,-0.2 3,-0.2 0.294 65.4 105.2 -98.0 7.0 11.5 -47.3 102.7 88 88 A N H > S+ 0 0 74 1,-0.2 4,-2.3 2,-0.2 -1,-0.2 0.837 75.9 51.6 -59.2 -37.3 11.7 -51.1 102.8 89 89 A L H > S+ 0 0 42 -3,-0.3 4,-1.7 2,-0.2 -1,-0.2 0.876 113.2 45.1 -69.4 -35.7 7.9 -51.7 102.7 90 90 A M H > S+ 0 0 0 -9,-0.3 4,-2.0 -3,-0.2 -2,-0.2 0.902 112.8 51.5 -72.1 -39.9 7.5 -49.4 99.7 91 91 A Q H X S+ 0 0 75 -4,-2.6 4,-2.2 1,-0.2 -2,-0.2 0.898 110.4 49.6 -63.0 -38.3 10.5 -51.0 98.0 92 92 A D H X S+ 0 0 57 -4,-2.3 4,-3.0 2,-0.2 5,-0.3 0.851 105.1 56.6 -69.0 -35.6 9.0 -54.4 98.7 93 93 A I H X S+ 0 0 0 -4,-1.7 4,-1.8 2,-0.2 -1,-0.2 0.946 110.5 45.5 -59.8 -47.0 5.6 -53.4 97.2 94 94 A L H X S+ 0 0 2 -4,-2.0 4,-2.2 1,-0.2 -2,-0.2 0.923 115.5 46.2 -61.8 -46.7 7.4 -52.5 94.0 95 95 A R H X S+ 0 0 102 -4,-2.2 4,-2.0 1,-0.2 -2,-0.2 0.870 113.4 46.8 -68.5 -36.3 9.4 -55.7 93.9 96 96 A E H X S+ 0 0 21 -4,-3.0 4,-2.3 2,-0.2 -1,-0.2 0.807 112.1 52.6 -74.3 -26.7 6.5 -58.0 94.7 97 97 A T H X S+ 0 0 0 -4,-1.8 4,-2.8 -5,-0.3 6,-0.3 0.891 107.8 51.6 -71.5 -39.1 4.5 -56.1 92.1 98 98 A E H X>S+ 0 0 10 -4,-2.2 4,-1.5 1,-0.2 5,-1.3 0.914 112.5 45.8 -61.7 -42.9 7.3 -56.7 89.6 99 99 A Q H <5S+ 0 0 119 -4,-2.0 -2,-0.2 2,-0.2 -1,-0.2 0.905 113.5 49.4 -67.6 -40.9 7.2 -60.4 90.5 100 100 A F H <5S+ 0 0 46 -4,-2.3 -2,-0.2 1,-0.2 -1,-0.2 0.924 121.1 33.1 -64.5 -46.8 3.4 -60.6 90.2 101 101 A L H <5S- 0 0 28 -4,-2.8 -1,-0.2 2,-0.2 -2,-0.2 0.631 101.1-129.0 -85.5 -13.7 3.2 -58.9 86.9 102 102 A N T <5 + 0 0 151 -4,-1.5 2,-0.4 1,-0.3 -3,-0.2 0.879 69.2 126.2 64.0 37.7 6.4 -60.3 85.5 103 103 A Q < - 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