==== Secondary Structure Definition by the program DSSP, updated CMBI version by ElmK / April 1,2000 ==== DATE=1-JAN-2010 . REFERENCE W. KABSCH AND C.SANDER, BIOPOLYMERS 22 (1983) 2577-2637 . HEADER GENE REGULATION/DNA 30-MAY-99 2IRF . COMPND 2 MOLECULE: DNA (5'-D(P*AP*AP*GP*TP*GP*AP*AP*AP*GP*(5IU) . SOURCE 2 SYNTHETIC: YES; . AUTHOR Y.FUJII,T.SHIMIZU,M.KUSUMOTO,Y.KYOGOKU,T.TANIGUCHI, . 654 6 0 0 0 TOTAL NUMBER OF RESIDUES, NUMBER OF CHAINS, NUMBER OF SS-BRIDGES(TOTAL,INTRACHAIN,INTERCHAIN) . 40843.0 ACCESSIBLE SURFACE OF PROTEIN (ANGSTROM**2) . 400 61.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(J) , SAME NUMBER PER 100 RESIDUES . 2 0.3 TOTAL NUMBER OF HYDROGEN BONDS IN PARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 101 15.4 TOTAL NUMBER OF HYDROGEN BONDS IN ANTIPARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 6 0.9 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-5), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-4), SAME NUMBER PER 100 RESIDUES . 1 0.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-3), SAME NUMBER PER 100 RESIDUES . 1 0.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-2), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-1), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+0), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+1), SAME NUMBER PER 100 RESIDUES . 26 4.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+2), SAME NUMBER PER 100 RESIDUES . 58 8.9 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+3), SAME NUMBER PER 100 RESIDUES . 189 28.9 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+4), SAME NUMBER PER 100 RESIDUES . 23 3.5 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+5), SAME NUMBER PER 100 RESIDUES . 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 *** HISTOGRAMS OF *** . 0 0 0 6 0 0 0 0 6 3 3 0 0 6 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 RESIDUES PER ALPHA HELIX . 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PARALLEL BRIDGES PER LADDER . 3 1 10 2 6 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 ANTIPARALLEL BRIDGES PER LADDER . 1 0 3 3 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 LADDERS PER SHEET . # RESIDUE AA STRUCTURE BP1 BP2 ACC N-H-->O O-->H-N N-H-->O O-->H-N TCO KAPPA ALPHA PHI PSI X-CA Y-CA Z-CA 1 5 G R 0 0 123 0, 0.0 2,-0.3 0, 0.0 57,-0.0 0.000 360.0 360.0 360.0 -26.8 41.1 102.6 89.6 2 6 G M - 0 0 78 55,-0.1 0, 0.0 1,-0.1 0, 0.0 -0.501 360.0-120.9 -67.1 122.7 38.1 104.7 90.9 3 7 G R > - 0 0 170 -2,-0.3 4,-2.1 1,-0.1 5,-0.2 -0.185 25.9-100.7 -61.8 160.7 36.0 105.8 87.9 4 8 G M H > S+ 0 0 8 1,-0.2 4,-2.4 2,-0.2 5,-0.1 0.830 113.0 51.4 -53.5 -46.3 35.5 109.5 87.2 5 9 G R H > S+ 0 0 35 2,-0.2 4,-2.4 1,-0.2 5,-0.2 0.970 114.6 39.3 -62.1 -57.6 31.9 109.9 88.6 6 10 G P H > S+ 0 0 55 0, 0.0 4,-1.9 0, 0.0 -1,-0.2 0.902 117.6 52.4 -60.1 -34.1 32.5 108.4 92.1 7 11 G W H X S+ 0 0 10 -4,-2.1 4,-1.4 1,-0.2 -2,-0.2 0.885 109.2 50.2 -64.6 -39.8 35.9 110.1 92.1 8 12 G L H X S+ 0 0 0 -4,-2.4 4,-2.5 -5,-0.2 -1,-0.2 0.881 107.0 52.6 -66.6 -43.1 34.3 113.4 91.2 9 13 G E H X S+ 0 0 42 -4,-2.4 4,-3.1 1,-0.2 5,-0.2 0.924 109.4 50.5 -59.3 -42.6 31.6 113.2 94.0 10 14 G E H X S+ 0 0 108 -4,-1.9 4,-1.2 1,-0.2 -1,-0.2 0.834 110.1 50.5 -63.1 -34.8 34.5 112.6 96.5 11 15 G Q H <>S+ 0 0 18 -4,-1.4 5,-0.7 2,-0.2 4,-0.5 0.897 113.8 44.0 -70.1 -42.2 36.3 115.7 95.1 12 16 G I H ><5S+ 0 0 1 -4,-2.5 3,-1.5 1,-0.2 -2,-0.2 0.950 113.7 50.0 -66.7 -49.6 33.1 117.8 95.5 13 17 G N H 3<5S+ 0 0 67 -4,-3.1 -1,-0.2 1,-0.3 -2,-0.2 0.730 103.2 61.3 -62.5 -24.7 32.3 116.5 98.9 14 18 G S T 3<5S- 0 0 47 -4,-1.2 -1,-0.3 -5,-0.2 -2,-0.2 0.652 91.0-146.4 -77.5 -17.6 35.9 117.1 100.1 15 19 G N T < 5 + 0 0 131 -3,-1.5 -3,-0.1 -4,-0.5 -2,-0.1 0.721 63.5 122.3 56.9 23.3 35.5 120.9 99.5 16 20 G T < + 0 0 91 -5,-0.7 -1,-0.1 1,-0.1 -4,-0.1 0.478 52.1 76.6 -91.7 -5.4 39.2 120.8 98.6 17 21 G I S > S- 0 0 7 -6,-0.4 3,-1.7 3,-0.3 2,-0.1 -0.931 86.8-123.7-111.0 115.1 38.7 122.3 95.1 18 22 G P T 3 S+ 0 0 67 0, 0.0 32,-0.1 0, 0.0 3,-0.1 -0.347 96.8 8.7 -58.1 120.5 38.0 126.0 95.0 19 23 G G T 3 S+ 0 0 37 1,-0.2 2,-1.0 -2,-0.1 12,-0.1 0.104 95.8 118.5 96.4 -23.6 34.7 126.6 93.2 20 24 G L < + 0 0 3 -3,-1.7 -3,-0.3 12,-0.1 2,-0.3 -0.691 41.3 128.9 -82.8 103.6 33.8 122.9 93.0 21 25 G K E -A 31 0A 109 10,-1.1 10,-2.5 -2,-1.0 2,-0.3 -0.955 61.0 -94.4-150.5 163.5 30.5 122.6 95.0 22 26 G W E -A 30 0A 42 -2,-0.3 8,-0.3 8,-0.2 3,-0.3 -0.636 19.4-166.5 -82.0 136.7 27.0 121.3 94.8 23 27 G L E S+ 0 0 57 6,-2.5 2,-0.2 1,-0.3 7,-0.2 0.747 94.2 3.6 -88.6 -29.9 24.4 123.8 93.6 24 28 G N E > -A 29 0A 60 5,-2.4 5,-2.1 -3,-0.1 -1,-0.3 -0.780 69.8-169.1-155.0 98.4 21.7 121.3 94.9 25 29 G K T > 5S+ 0 0 131 -3,-0.3 3,-0.5 -2,-0.2 -1,-0.1 0.845 86.2 58.0 -61.2 -35.6 23.1 118.3 96.6 26 30 G E T 3 5S+ 0 0 164 1,-0.2 -1,-0.2 2,-0.1 -2,-0.0 0.960 113.9 35.0 -59.5 -55.7 19.7 116.5 96.6 27 31 G K T 3 5S- 0 0 136 2,-0.1 -1,-0.2 1,-0.0 80,-0.2 0.362 112.0-121.7 -82.0 5.4 19.3 116.7 92.8 28 32 G K T < 5 + 0 0 57 -3,-0.5 79,-1.8 -4,-0.3 2,-0.5 0.770 59.2 151.6 58.1 31.0 23.0 116.2 92.4 29 33 G I E < +AB 24 106A 19 -5,-2.1 -6,-2.5 77,-0.2 -5,-2.4 -0.807 16.1 162.7 -91.9 127.2 23.4 119.4 90.4 30 34 G F E -AB 22 105A 0 75,-2.8 75,-2.7 -2,-0.5 2,-0.4 -0.856 29.7-127.2-138.0 172.2 26.9 120.9 90.9 31 35 G Q E -AB 21 104A 31 -10,-2.5 -10,-1.1 -2,-0.3 73,-0.2 -0.967 8.7-154.2-128.8 144.6 29.2 123.4 89.3 32 36 G I E - B 0 103A 0 71,-2.8 71,-2.6 -2,-0.4 2,-0.2 -0.955 28.3-116.1-115.3 131.7 32.8 123.1 88.1 33 37 G P E + B 0 102A 18 0, 0.0 69,-0.3 0, 0.0 68,-0.1 -0.512 35.1 178.1 -66.6 135.9 35.0 126.3 88.0 34 38 G W + 0 0 15 67,-3.0 2,-0.3 -2,-0.2 68,-0.2 -0.281 7.0 165.8-138.6 52.6 36.0 126.9 84.3 35 39 G M - 0 0 5 66,-0.3 64,-0.1 12,-0.1 2,-0.1 -0.533 42.8-110.2 -75.5 134.7 38.1 130.0 84.1 36 40 G H > - 0 0 114 62,-0.4 3,-0.9 -2,-0.3 6,-0.1 -0.368 17.9-137.0 -62.7 133.6 39.9 130.5 80.8 37 41 G A T 3 S+ 0 0 49 1,-0.2 -1,-0.1 -2,-0.1 4,-0.1 0.509 102.8 56.4 -71.9 -6.3 43.7 130.1 81.1 38 42 G A T 3 S+ 0 0 84 2,-0.1 2,-0.3 3,-0.0 -1,-0.2 0.618 87.2 98.1 -97.0 -18.4 44.3 133.2 78.9 39 43 G R S X S- 0 0 109 -3,-0.9 3,-1.8 1,-0.1 -4,-0.0 -0.543 85.1-106.4 -77.0 134.0 42.2 135.4 81.2 40 44 G H T 3 S+ 0 0 178 -2,-0.3 -1,-0.1 1,-0.3 3,-0.1 -0.305 107.0 41.1 -57.0 132.2 44.0 137.7 83.7 41 45 G G T 3 S+ 0 0 35 1,-0.5 -1,-0.3 -4,-0.1 2,-0.2 0.184 81.8 131.5 110.5 -15.7 43.4 136.3 87.2 42 46 G W < + 0 0 71 -3,-1.8 -1,-0.5 -6,-0.1 2,-0.3 -0.496 30.8 178.2 -71.7 141.8 43.9 132.7 86.2 43 47 G D >> - 0 0 69 -2,-0.2 4,-3.1 1,-0.1 5,-0.6 -0.994 34.6-132.9-146.5 136.9 46.2 130.7 88.4 44 48 G V H >5S+ 0 0 37 -2,-0.3 4,-1.7 1,-0.2 8,-0.2 0.912 108.6 48.0 -58.5 -41.1 47.2 127.0 88.1 45 49 G E H 45S+ 0 0 108 1,-0.2 -1,-0.2 3,-0.2 -3,-0.0 0.793 120.0 36.9 -70.4 -31.9 46.7 126.5 91.9 46 50 G K H 45S+ 0 0 78 -3,-0.1 -2,-0.2 2,-0.1 -1,-0.2 0.846 130.0 26.0 -90.0 -37.0 43.3 128.2 92.0 47 51 G D H <5S+ 0 0 12 -4,-3.1 -3,-0.2 1,-0.1 4,-0.2 0.693 123.2 41.3-104.7 -21.1 41.8 127.2 88.6 48 52 G A S X S+ 0 0 12 0, 0.0 4,-2.5 0, 0.0 5,-0.1 0.867 85.9 49.4 -54.0 -38.0 43.6 121.7 90.9 50 54 G L H > S+ 0 0 2 2,-0.2 4,-1.5 1,-0.2 5,-0.1 0.884 113.3 45.8 -67.1 -39.7 40.7 119.6 89.8 51 55 G F H > S+ 0 0 7 -3,-0.4 4,-1.7 2,-0.2 -1,-0.2 0.884 113.8 49.3 -70.3 -38.4 42.2 118.9 86.4 52 56 G R H X S+ 0 0 46 -4,-2.8 4,-2.5 1,-0.2 -2,-0.2 0.919 108.3 54.2 -65.6 -44.1 45.6 118.2 87.9 53 57 G N H X S+ 0 0 47 -4,-2.5 4,-2.6 -5,-0.3 -1,-0.2 0.845 108.0 48.8 -58.5 -40.1 44.0 115.8 90.4 54 58 G W H X S+ 0 0 32 -4,-1.5 4,-1.8 2,-0.2 6,-0.3 0.894 111.5 50.4 -68.1 -38.1 42.4 113.8 87.6 55 59 G A H <>S+ 0 0 6 -4,-1.7 5,-2.0 2,-0.2 6,-1.5 0.867 113.7 44.3 -66.7 -38.5 45.7 113.7 85.8 56 60 G I H ><5S+ 0 0 65 -4,-2.5 3,-1.6 4,-0.2 -2,-0.2 0.938 112.4 52.7 -70.7 -44.7 47.5 112.5 88.9 57 61 G H H 3<5S+ 0 0 63 -4,-2.6 -2,-0.2 1,-0.3 -1,-0.2 0.798 114.2 42.7 -59.8 -33.4 44.7 110.0 89.6 58 62 G T T 3<5S- 0 0 52 -4,-1.8 -1,-0.3 -5,-0.1 -2,-0.2 0.235 115.4-111.5-100.2 11.5 45.0 108.6 86.1 59 63 G G T < 5S+ 0 0 56 -3,-1.6 -3,-0.2 2,-0.2 -4,-0.1 0.550 86.4 117.3 72.6 9.4 48.8 108.5 85.9 60 64 G K S > - 0 0 101 -2,-0.4 4,-2.3 -3,-0.1 3,-1.6 -0.935 33.1-122.9-115.8 103.6 53.6 112.8 88.2 63 67 G P T 34 S+ 0 0 88 0, 0.0 4,-0.1 0, 0.0 2,-0.1 -0.187 95.8 9.5 -48.0 124.8 54.2 115.9 90.4 64 68 G G T 34 S+ 0 0 85 2,-0.5 3,-0.0 1,-0.1 -3,-0.0 -0.113 128.0 59.9 95.0 -38.0 57.9 116.8 90.1 65 69 G I T <4 S+ 0 0 121 -3,-1.6 2,-0.3 1,-0.1 -1,-0.1 0.935 98.7 52.9 -86.1 -58.0 58.6 114.5 87.2 66 70 G D S < S- 0 0 64 -4,-2.3 -2,-0.5 1,-0.1 -1,-0.1 -0.587 88.9-115.0 -83.1 141.5 56.2 115.7 84.5 67 71 G K - 0 0 175 -2,-0.3 -1,-0.1 1,-0.1 -5,-0.0 -0.546 45.8 -93.5 -73.0 134.0 56.1 119.4 83.5 68 72 G P - 0 0 71 0, 0.0 -1,-0.1 0, 0.0 -7,-0.0 -0.243 43.7-179.3 -51.5 129.8 52.8 121.0 84.4 69 73 G D >> + 0 0 73 1,-0.1 4,-2.3 -3,-0.1 3,-0.7 -0.528 10.5 166.6-134.9 68.5 50.4 120.9 81.4 70 74 G P H 3> S+ 0 0 29 0, 0.0 4,-2.4 0, 0.0 5,-0.2 0.788 76.0 59.4 -50.6 -39.4 47.1 122.6 82.5 71 75 G K H 3> S+ 0 0 165 2,-0.2 4,-2.1 1,-0.2 5,-0.1 0.909 110.9 39.4 -60.5 -45.0 45.8 122.8 78.9 72 76 G T H <> S+ 0 0 60 -3,-0.7 4,-3.3 2,-0.2 5,-0.3 0.895 112.5 56.2 -73.0 -40.8 45.9 119.0 78.5 73 77 G W H X S+ 0 0 12 -4,-2.3 4,-1.8 1,-0.2 -2,-0.2 0.912 113.0 42.7 -56.3 -43.6 44.7 118.4 82.1 74 78 G K H X S+ 0 0 44 -4,-2.4 4,-2.6 2,-0.2 -2,-0.2 0.921 115.3 47.2 -69.8 -45.4 41.6 120.5 81.3 75 79 G A H X S+ 0 0 33 -4,-2.1 4,-2.5 1,-0.2 5,-0.2 0.911 113.7 49.0 -63.5 -41.9 41.0 119.0 77.8 76 80 G N H X S+ 0 0 77 -4,-3.3 4,-2.3 1,-0.2 -1,-0.2 0.896 112.5 48.3 -63.2 -42.5 41.4 115.5 79.1 77 81 G F H X S+ 0 0 0 -4,-1.8 4,-2.4 -5,-0.3 -2,-0.2 0.912 112.0 48.5 -65.1 -46.1 39.0 116.2 81.9 78 82 G R H X S+ 0 0 80 -4,-2.6 4,-2.6 2,-0.2 -2,-0.2 0.925 112.4 47.6 -61.6 -47.6 36.4 117.8 79.6 79 83 G C H X S+ 0 0 70 -4,-2.5 4,-2.0 2,-0.2 -2,-0.2 0.935 112.2 50.1 -60.2 -45.0 36.5 114.9 77.1 80 84 G A H X S+ 0 0 21 -4,-2.3 4,-0.8 1,-0.2 -1,-0.2 0.899 111.3 49.5 -59.6 -41.2 36.2 112.4 80.0 81 85 G M H >< S+ 0 0 2 -4,-2.4 3,-0.8 1,-0.2 -1,-0.2 0.901 107.7 54.2 -63.6 -43.4 33.2 114.4 81.3 82 86 G N H 3< S+ 0 0 99 -4,-2.6 -1,-0.2 1,-0.2 -2,-0.2 0.871 107.2 51.8 -58.6 -39.9 31.6 114.4 77.8 83 87 G S H 3< S+ 0 0 94 -4,-2.0 -1,-0.2 -5,-0.2 -2,-0.2 0.649 82.7 107.0 -74.8 -17.1 31.9 110.6 77.6 84 88 G L X< - 0 0 35 -4,-0.8 3,-0.6 -3,-0.8 -3,-0.0 -0.447 55.5-158.1 -67.4 128.0 30.2 109.9 81.0 85 89 G P T 3 S+ 0 0 108 0, 0.0 -1,-0.1 0, 0.0 -4,-0.0 0.753 89.3 50.0 -77.5 -22.1 26.7 108.4 80.5 86 90 G D T 3 S+ 0 0 25 1,-0.1 22,-2.0 21,-0.1 2,-0.3 0.145 103.5 60.0-104.2 16.3 25.6 109.5 84.0 87 91 G I E < +C 107 0A 8 -3,-0.6 2,-0.3 20,-0.3 20,-0.2 -0.958 60.3 174.6-143.9 123.7 26.6 113.1 84.1 88 92 G E E -C 106 0A 89 18,-2.1 18,-2.5 -2,-0.3 2,-0.2 -0.946 31.6-108.1-130.5 152.7 25.4 115.7 81.7 89 93 G E E -C 105 0A 33 -2,-0.3 2,-0.9 16,-0.2 16,-0.3 -0.513 12.9-145.7 -80.9 143.4 25.9 119.5 81.4 90 94 G V >> - 0 0 22 14,-3.0 4,-1.4 -2,-0.2 3,-1.4 -0.828 23.7-179.3-106.3 82.3 23.1 121.9 82.2 91 95 G K T 34 S+ 0 0 147 -2,-0.9 -1,-0.2 1,-0.2 14,-0.0 0.759 71.2 66.1 -57.2 -30.9 24.1 124.4 79.6 92 96 G D T 34 S+ 0 0 142 1,-0.2 -1,-0.2 -3,-0.1 -2,-0.0 0.768 111.5 36.6 -65.7 -22.8 21.3 126.9 80.4 93 97 G R T <4 S+ 0 0 105 -3,-1.4 -1,-0.2 11,-0.1 -2,-0.2 0.710 96.8 99.5 -99.3 -26.2 22.9 127.5 83.8 94 98 G S < - 0 0 20 -4,-1.4 9,-0.2 10,-0.2 2,-0.1 -0.266 45.5-175.0 -63.5 146.6 26.6 127.3 82.7 95 99 G I B +D 102 0A 95 7,-2.4 7,-1.8 5,-0.1 -61,-0.2 -0.615 10.8 168.2-143.9 75.8 28.6 130.5 82.2 96 100 G K + 0 0 117 5,-0.3 2,-0.3 -2,-0.1 5,-0.1 0.129 55.5 82.3 -79.7 22.4 32.0 129.5 80.8 97 101 G K S S+ 0 0 162 5,-0.2 5,-0.2 2,-0.1 -63,-0.1 -0.807 77.2 15.3-122.5 162.9 32.8 133.1 79.9 98 102 G G S > S- 0 0 28 -2,-0.3 3,-0.7 3,-0.1 -62,-0.4 -0.095 103.8 -22.7 71.7-172.1 34.2 136.1 81.8 99 103 G N T 3 S+ 0 0 91 1,-0.2 -2,-0.1 -64,-0.1 -58,-0.1 -0.503 131.2 12.3 -75.5 142.3 35.8 136.2 85.3 100 104 G N T 3 S- 0 0 109 -2,-0.2 -1,-0.2 1,-0.1 -3,-0.1 0.904 89.3-174.5 55.5 46.5 35.1 133.2 87.6 101 105 G A < + 0 0 4 -3,-0.7 -67,-3.0 -5,-0.1 2,-0.3 -0.309 5.4 174.8 -67.7 153.4 33.6 131.3 84.7 102 106 G F E -BD 33 95A 45 -7,-1.8 -7,-2.4 -69,-0.3 2,-0.3 -0.976 25.8-134.5-156.6 165.6 31.9 127.9 85.6 103 107 G R E -B 32 0A 20 -71,-2.6 -71,-2.8 -2,-0.3 2,-0.4 -0.935 17.6-140.6-125.8 146.0 29.9 125.0 84.1 104 108 G V E +B 31 0A 0 -2,-0.3 -14,-3.0 -14,-0.3 2,-0.3 -0.932 22.1 178.8-111.3 134.9 26.9 123.3 85.7 105 109 G Y E -BC 30 89A 4 -75,-2.7 -75,-2.8 -2,-0.4 2,-0.4 -0.931 17.0-150.4-131.5 156.0 26.4 119.6 85.6 106 110 G R E -BC 29 88A 75 -18,-2.5 -18,-2.1 -2,-0.3 2,-0.6 -0.996 17.5-133.6-128.7 131.6 23.8 117.2 86.9 107 111 G M E - 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