==== Secondary Structure Definition by the program DSSP, updated CMBI version by ElmK / April 1,2000 ==== DATE=5-DEC-2009 . REFERENCE W. KABSCH AND C.SANDER, BIOPOLYMERS 22 (1983) 2577-2637 . HEADER HYDROLASE 08-JAN-03 1J2P . COMPND 2 MOLECULE: PROTEASOME ALPHA SUBUNIT; . SOURCE 2 ORGANISM_SCIENTIFIC: ARCHAEOGLOBUS FULGIDUS; . AUTHOR M.GROLL,H.BRANDSTETTER,H.BARTUNIK,G.BOURENKOW,R.HUBER . 1701 7 0 0 0 TOTAL NUMBER OF RESIDUES, NUMBER OF CHAINS, NUMBER OF SS-BRIDGES(TOTAL,INTRACHAIN,INTERCHAIN) . 71793.0 ACCESSIBLE SURFACE OF PROTEIN (ANGSTROM**2) . 1175 69.1 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(J) , SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS IN PARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 343 20.2 TOTAL NUMBER OF HYDROGEN BONDS IN ANTIPARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-5), SAME NUMBER PER 100 RESIDUES . 21 1.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-4), SAME NUMBER PER 100 RESIDUES . 14 0.8 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-3), SAME NUMBER PER 100 RESIDUES . 7 0.4 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-2), SAME NUMBER PER 100 RESIDUES . 14 0.8 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-1), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+0), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+1), SAME NUMBER PER 100 RESIDUES . 62 3.6 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+2), SAME NUMBER PER 100 RESIDUES . 154 9.1 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+3), SAME NUMBER PER 100 RESIDUES . 546 32.1 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+4), SAME NUMBER PER 100 RESIDUES . 7 0.4 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+5), SAME NUMBER PER 100 RESIDUES . 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 *** HISTOGRAMS OF *** . 0 0 0 0 0 0 0 0 0 0 14 0 0 7 0 7 0 7 0 0 7 0 0 0 0 0 0 0 0 0 RESIDUES PER ALPHA HELIX . 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PARALLEL BRIDGES PER LADDER . 14 7 14 7 7 7 14 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 ANTIPARALLEL BRIDGES PER LADDER . 0 0 0 14 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 LADDERS PER SHEET . # RESIDUE AA STRUCTURE BP1 BP2 ACC N-H-->O O-->H-N N-H-->O O-->H-N TCO KAPPA ALPHA PHI PSI X-CA Y-CA Z-CA 1 4 A P 0 0 89 0, 0.0 2,-0.5 0, 0.0 5,-0.0 0.000 360.0 360.0 360.0 56.8 46.1 94.7 -5.8 2 5 A Q > - 0 0 119 1,-0.0 3,-1.6 3,-0.0 4,-0.4 -0.924 360.0-124.7-113.3 122.4 48.2 91.8 -7.1 3 6 A M T 3 S+ 0 0 111 -2,-0.5 1465,-0.0 1,-0.2 -1,-0.0 -0.154 95.1 46.9 -54.6 153.1 47.1 88.2 -6.7 4 7 A G T 3 S+ 0 0 22 9,-0.0 -1,-0.2 243,-0.0 3,-0.0 0.027 116.9 44.9 99.1 -27.7 49.7 86.0 -5.0 5 8 A Y S < S+ 0 0 29 -3,-1.6 14,-0.4 1,-0.0 3,-0.3 0.355 97.3 72.3-127.6 2.6 50.2 88.5 -2.2 6 9 A D S S+ 0 0 1 -4,-0.4 -3,-0.1 1,-0.2 3,-0.1 0.223 79.6 65.6-110.0 17.0 46.7 89.5 -1.2 7 10 A R S S+ 0 0 123 1,-0.2 2,-0.3 1468,-0.1 -1,-0.2 0.132 90.2 61.8-125.4 27.4 45.0 86.6 0.5 8 11 A A > - 0 0 42 -3,-0.3 3,-2.6 3,-0.2 12,-0.2 -0.860 66.5-147.6-152.0 110.2 47.0 86.3 3.8 9 12 A I T 3 S+ 0 0 10 -2,-0.3 12,-0.1 1,-0.3 115,-0.1 0.742 98.0 57.3 -49.1 -29.2 47.2 89.1 6.3 10 13 A T T 3 S+ 0 0 25 114,-0.1 2,-0.5 -3,-0.1 -1,-0.3 0.489 87.9 93.3 -85.9 -1.9 50.7 88.2 7.3 11 14 A V < - 0 0 14 -3,-2.6 8,-0.3 -6,-0.1 2,-0.3 -0.801 62.6-148.9-100.4 129.3 52.2 88.5 3.8 12 15 A F - 0 0 0 -2,-0.5 6,-0.2 6,-0.1 256,-0.1 -0.647 19.6-127.8 -86.2 148.7 53.8 91.7 2.6 13 16 A S > - 0 0 0 4,-1.6 3,-2.1 -2,-0.3 4,-0.1 -0.440 34.3 -95.7 -88.1 169.3 53.6 92.4 -1.1 14 17 A P T 3 S+ 0 0 52 0, 0.0 -1,-0.1 0, 0.0 256,-0.1 0.809 127.9 50.5 -58.7 -27.2 56.7 93.2 -3.2 15 18 A D T 3 S- 0 0 72 2,-0.0 252,-0.0 255,-0.0 253,-0.0 0.046 126.2 -98.7 -99.1 28.4 56.1 96.9 -2.7 16 19 A G S < S+ 0 0 0 -3,-2.1 2,-0.3 1,-0.2 258,-0.1 0.879 75.0 134.5 63.2 47.6 55.6 96.6 1.1 17 20 A R - 0 0 68 -4,-0.1 -4,-1.6 -12,-0.0 2,-0.6 -0.831 54.3-138.2-126.3 163.2 51.9 96.5 1.6 18 21 A L >> - 0 0 13 -2,-0.3 4,-1.7 -6,-0.2 3,-1.5 -0.808 22.3-177.5-116.8 81.4 49.3 94.5 3.5 19 22 A F H 3> S+ 0 0 5 -2,-0.6 4,-3.0 -14,-0.4 5,-0.2 0.814 78.7 61.8 -52.9 -34.4 46.6 94.0 0.9 20 23 A Q H 3> S+ 0 0 5 -15,-0.3 4,-1.9 -9,-0.2 -1,-0.3 0.837 106.0 48.2 -64.0 -28.0 44.2 92.2 3.3 21 24 A V H <> S+ 0 0 4 -3,-1.5 4,-2.0 2,-0.2 -1,-0.2 0.892 111.4 49.1 -75.1 -40.5 44.2 95.4 5.4 22 25 A E H X S+ 0 0 46 -4,-1.7 4,-0.8 1,-0.2 -2,-0.2 0.892 112.1 49.2 -65.8 -36.2 43.5 97.5 2.3 23 26 A Y H >X S+ 0 0 14 -4,-3.0 3,-1.0 1,-0.2 4,-0.7 0.905 106.6 54.9 -68.7 -40.6 40.7 95.1 1.4 24 27 A A H >X S+ 0 0 0 -4,-1.9 3,-1.5 1,-0.3 4,-1.0 0.900 102.3 58.7 -58.8 -38.2 39.3 95.4 5.0 25 28 A R H 3X S+ 0 0 96 -4,-2.0 4,-1.3 1,-0.3 -1,-0.3 0.793 94.3 65.0 -61.5 -26.0 39.3 99.1 4.4 26 29 A E H < S+ 0 0 13 -4,-1.0 3,-1.5 1,-0.2 49,-0.2 0.817 99.7 67.5 -69.6 -24.9 34.3 100.4 5.9 29 32 A K H 3< S+ 0 0 113 -4,-1.3 -1,-0.2 1,-0.3 -2,-0.2 0.830 95.5 57.5 -59.8 -32.8 33.8 102.4 2.7 30 33 A R T << S+ 0 0 124 -3,-0.8 -1,-0.3 -4,-0.7 -2,-0.2 0.546 88.7 101.2 -76.3 -10.1 30.3 100.8 2.3 31 34 A G S < S- 0 0 14 -3,-1.5 132,-2.4 -4,-0.3 3,-0.1 -0.351 87.5 -87.6 -80.0 159.0 29.0 101.9 5.7 32 35 A A - 0 0 25 130,-0.2 2,-0.2 45,-0.1 -1,-0.1 -0.342 60.0-100.7 -57.0 141.3 26.7 104.8 6.4 33 36 A T - 0 0 0 129,-0.1 15,-1.8 -3,-0.1 2,-0.4 -0.438 35.6-163.9 -74.7 139.5 28.8 107.9 6.8 34 37 A A E -AB 47 161A 2 127,-2.2 127,-2.1 13,-0.2 2,-0.3 -0.985 7.1-155.4-123.6 131.8 29.7 109.2 10.3 35 38 A I E -AB 46 160A 0 11,-4.0 11,-2.8 -2,-0.4 2,-0.4 -0.827 7.2-167.0-111.5 146.7 31.0 112.7 10.8 36 39 A G E +AB 45 159A 0 123,-2.4 123,-2.7 -2,-0.3 2,-0.4 -0.998 9.4 174.2-133.0 130.6 33.1 114.1 13.6 37 40 A I E -AB 44 158A 0 7,-2.5 7,-3.1 -2,-0.4 2,-0.3 -0.992 10.5-161.4-141.4 131.5 33.8 117.8 14.2 38 41 A K E +A 43 0A 19 119,-3.4 2,-0.2 -2,-0.4 5,-0.2 -0.808 14.3 166.1-111.4 151.5 35.6 119.5 17.1 39 42 A C E > -A 42 0A 13 3,-2.6 3,-0.6 -2,-0.3 142,-0.1 -0.827 51.7 -77.0-146.9-174.8 35.5 123.1 18.3 40 43 A K T 3 S+ 0 0 136 140,-0.9 141,-0.1 -2,-0.2 3,-0.1 0.914 125.3 36.4 -56.5 -51.2 36.4 125.3 21.2 41 44 A E T 3 S- 0 0 72 1,-0.1 172,-2.6 171,-0.1 173,-0.4 0.430 126.0 -44.1 -88.1 -0.3 33.6 124.2 23.5 42 45 A G E < -AC 39 212A 0 -3,-0.6 -3,-2.6 170,-0.3 2,-0.4 -0.990 57.0 -78.4 164.6-173.2 33.3 120.5 22.6 43 46 A V E -AC 38 211A 1 168,-2.2 168,-2.1 -2,-0.3 2,-0.4 -0.957 31.5-157.9-125.9 142.6 33.2 117.7 20.1 44 47 A I E -AC 37 210A 0 -7,-3.1 -7,-2.5 -2,-0.4 2,-0.4 -0.925 5.5-163.0-124.1 146.1 30.3 116.5 17.9 45 48 A L E -AC 36 209A 0 164,-2.7 164,-2.2 -2,-0.4 2,-0.4 -0.993 8.9-174.5-128.2 124.8 29.4 113.2 16.2 46 49 A I E -AC 35 208A 1 -11,-2.8 -11,-4.0 -2,-0.4 2,-0.4 -0.980 4.7-177.2-123.9 128.9 26.9 113.0 13.4 47 50 A A E -AC 34 207A 2 160,-2.8 160,-3.1 -2,-0.4 2,-0.5 -0.953 26.7-124.5-128.9 145.7 25.7 109.8 11.8 48 51 A D E + C 0 206A 6 -15,-1.8 158,-0.2 -2,-0.4 -16,-0.1 -0.749 27.5 174.1 -86.4 122.6 23.4 108.9 9.0 49 52 A K + 0 0 44 156,-1.2 157,-0.2 -2,-0.5 -1,-0.1 0.117 22.1 143.7-117.8 22.7 20.7 106.5 10.1 50 53 A R - 0 0 121 155,-0.5 2,-0.3 2,-0.0 155,-0.1 -0.408 34.7-155.9 -62.6 132.9 18.5 106.3 7.0 51 54 A V - 0 0 53 -2,-0.1 -2,-0.1 1,-0.1 -1,-0.0 -0.827 20.8-136.8-113.2 153.4 17.1 102.8 6.6 52 55 A G S S+ 0 0 89 -2,-0.3 2,-0.3 1,-0.1 -1,-0.1 0.526 86.6 47.1 -84.6 -5.1 15.9 101.1 3.5 53 56 A S > - 0 0 20 1,-0.1 3,-0.6 3,-0.0 -1,-0.1 -0.989 62.3-146.5-140.6 148.2 12.9 99.5 5.2 54 57 A K T 3 S+ 0 0 96 -2,-0.3 4,-0.1 1,-0.2 -1,-0.1 0.506 95.1 75.1 -85.4 -7.0 10.1 100.6 7.5 55 58 A L T 3 S+ 0 0 1 1566,-0.1 1566,-2.8 1565,-0.1 1567,-0.7 0.678 74.6 95.4 -78.6 -17.6 10.1 97.2 9.1 56 59 A L B < S-E 1620 0B 24 -3,-0.6 2,-2.8 1564,-0.3 1564,-0.3 -0.569 85.6-123.9 -75.8 134.4 13.3 98.0 11.0 57 60 A E > + 0 0 26 1562,-3.2 3,-2.1 -2,-0.3 1562,-0.1 -0.459 43.6 167.3 -75.3 70.7 12.7 99.3 14.5 58 61 A K G > + 0 0 85 -2,-2.8 3,-1.5 1,-0.3 -1,-0.2 0.696 63.4 78.7 -62.3 -17.4 14.6 102.5 13.7 59 62 A D G 3 S+ 0 0 135 1,-0.3 -1,-0.3 -3,-0.2 -2,-0.1 0.576 104.3 36.4 -66.5 -6.5 13.3 104.0 16.9 60 63 A T G < S+ 0 0 50 -3,-2.1 2,-0.3 1559,-0.1 -1,-0.3 0.195 87.3 123.3-129.1 11.0 15.9 101.9 18.5 61 64 A I < - 0 0 46 -3,-1.5 2,-0.5 -5,-0.1 1557,-0.0 -0.599 42.9-162.6 -77.7 135.3 18.7 102.2 16.0 62 65 A E + 0 0 79 -2,-0.3 3,-0.1 1,-0.1 -3,-0.0 -0.974 27.0 160.6-126.1 122.8 22.0 103.5 17.4 63 66 A K S S+ 0 0 62 -2,-0.5 12,-0.7 1,-0.2 2,-0.5 0.601 72.6 55.5-109.1 -21.5 24.8 104.9 15.3 64 67 A I E S-F 74 0C 1 10,-0.2 2,-0.4 -17,-0.1 10,-0.2 -0.966 75.6-159.5-116.2 123.9 26.5 106.9 18.0 65 68 A Y E -F 73 0C 49 8,-3.5 8,-1.1 -2,-0.5 2,-0.4 -0.884 15.6-133.0-114.7 136.1 27.6 105.0 21.2 66 69 A K E +F 72 0C 74 -2,-0.4 6,-0.2 6,-0.2 3,-0.1 -0.652 23.4 175.7 -79.5 129.5 28.3 106.1 24.7 67 70 A I - 0 0 12 4,-2.4 2,-0.3 1,-0.5 5,-0.2 0.771 69.2 -20.7-100.8 -41.1 31.5 104.5 25.9 68 71 A D S S- 0 0 19 3,-1.4 -1,-0.5 25,-0.0 3,-0.2 -0.876 86.4 -75.3-154.4 179.3 31.8 106.3 29.2 69 72 A E S S+ 0 0 92 -2,-0.3 149,-3.5 1,-0.2 -3,-0.0 0.849 133.0 23.2 -54.9 -35.3 30.4 109.5 30.7 70 73 A H S S+ 0 0 18 147,-0.2 68,-1.0 141,-0.1 2,-0.4 0.442 108.8 86.0-111.4 -4.2 32.8 111.6 28.6 71 74 A I E - G 0 137C 0 -3,-0.2 -4,-2.4 66,-0.2 -3,-1.4 -0.865 51.4-177.1-108.5 139.8 33.7 109.3 25.8 72 75 A C E -FG 66 136C 1 64,-4.7 64,-2.5 -2,-0.4 2,-0.3 -0.833 1.9-169.6-127.3 158.6 31.7 109.0 22.6 73 76 A A E -FG 65 135C 1 -8,-1.1 -8,-3.5 -2,-0.3 2,-0.3 -0.966 11.8-167.2-160.3 141.8 32.0 106.8 19.4 74 77 A A E -FG 64 134C 1 60,-2.1 60,-1.5 -2,-0.3 2,-0.3 -0.793 18.5-146.6-119.2 165.4 30.7 106.2 15.9 75 78 A T E + G 0 133C 17 -12,-0.7 2,-0.3 -2,-0.3 58,-0.2 -0.926 17.7 175.6-138.0 156.0 31.2 103.2 13.5 76 79 A S E + G 0 132C 3 56,-1.2 56,-2.3 -2,-0.3 2,-0.3 -0.906 50.6 32.4-160.2 131.5 31.6 102.1 9.9 77 80 A G E S+ G 0 131C 19 -2,-0.3 54,-0.2 54,-0.2 -45,-0.1 -0.903 98.7 18.2 125.3-155.2 32.2 98.7 8.3 78 81 A L > - 0 0 33 52,-2.6 4,-2.2 -2,-0.3 5,-0.1 -0.327 67.3-142.0 -55.5 125.4 31.2 95.2 9.2 79 82 A V H > S+ 0 0 85 2,-0.2 4,-2.1 1,-0.2 5,-0.2 0.932 95.3 43.1 -56.2 -59.7 28.3 95.3 11.6 80 83 A A H > S+ 0 0 3 1,-0.2 4,-2.0 2,-0.2 -1,-0.2 0.895 113.8 53.3 -56.7 -43.3 29.1 92.5 14.0 81 84 A D H > S+ 0 0 2 1,-0.2 4,-1.9 2,-0.2 -1,-0.2 0.922 108.5 51.2 -58.0 -44.6 32.8 93.5 14.1 82 85 A A H X S+ 0 0 4 -4,-2.2 4,-2.2 1,-0.2 -2,-0.2 0.931 107.1 51.0 -59.7 -50.0 31.8 97.0 15.1 83 86 A R H X S+ 0 0 38 -4,-2.1 4,-2.2 1,-0.2 5,-0.2 0.911 109.6 51.6 -54.5 -45.2 29.6 96.0 18.0 84 87 A V H X S+ 0 0 26 -4,-2.0 4,-1.9 1,-0.2 -1,-0.2 0.883 108.5 51.2 -60.8 -40.1 32.4 93.8 19.4 85 88 A L H X S+ 0 0 0 -4,-1.9 4,-1.9 1,-0.2 -1,-0.2 0.885 109.7 49.6 -65.9 -40.0 34.8 96.7 19.2 86 89 A I H X S+ 0 0 7 -4,-2.2 4,-2.3 2,-0.2 -2,-0.2 0.889 110.0 49.0 -68.6 -38.3 32.4 99.0 21.1 87 90 A D H X S+ 0 0 29 -4,-2.2 4,-2.8 1,-0.2 5,-0.2 0.905 110.0 54.6 -65.3 -37.8 31.8 96.5 23.9 88 91 A R H X S+ 0 0 100 -4,-1.9 4,-2.4 -5,-0.2 -2,-0.2 0.899 107.2 49.5 -60.5 -41.4 35.6 96.2 24.0 89 92 A A H X S+ 0 0 4 -4,-1.9 4,-2.0 2,-0.2 -1,-0.2 0.920 111.2 49.0 -64.8 -44.7 35.9 99.9 24.5 90 93 A R H X S+ 0 0 65 -4,-2.3 4,-1.9 2,-0.2 -2,-0.2 0.945 111.7 48.0 -60.6 -48.6 33.3 100.0 27.2 91 94 A I H X S+ 0 0 92 -4,-2.8 4,-2.8 1,-0.2 -1,-0.2 0.930 111.5 52.2 -56.7 -44.0 35.0 97.1 29.1 92 95 A E H X S+ 0 0 31 -4,-2.4 4,-1.5 1,-0.2 -1,-0.2 0.850 106.8 52.8 -61.2 -35.5 38.3 98.9 28.6 93 96 A A H X S+ 0 0 2 -4,-2.0 4,-1.2 2,-0.2 11,-0.3 0.852 111.3 46.5 -68.9 -34.8 36.7 102.1 30.2 94 97 A Q H X S+ 0 0 121 -4,-1.9 4,-1.3 -3,-0.2 -2,-0.2 0.910 107.0 56.1 -73.3 -44.2 35.6 100.1 33.2 95 98 A I H X S+ 0 0 72 -4,-2.8 4,-2.8 1,-0.2 3,-0.4 0.894 105.1 55.2 -54.2 -41.1 38.9 98.3 33.7 96 99 A N H X S+ 0 0 11 -4,-1.5 4,-2.7 1,-0.2 6,-0.8 0.908 104.2 50.4 -60.3 -46.1 40.5 101.8 33.9 97 100 A R H < S+ 0 0 101 -4,-1.2 -1,-0.2 4,-0.2 -2,-0.2 0.751 115.3 46.6 -65.0 -22.8 38.3 103.1 36.7 98 101 A L H < S+ 0 0 127 -4,-1.3 -2,-0.2 -3,-0.4 -1,-0.2 0.923 118.3 37.5 -82.4 -51.6 39.2 99.9 38.6 99 102 A T H < S+ 0 0 83 -4,-2.8 -2,-0.2 1,-0.2 -3,-0.2 0.888 136.5 19.1 -69.1 -46.2 42.9 99.8 38.0 100 103 A Y S < S- 0 0 134 -4,-2.7 -3,-0.2 2,-0.3 -1,-0.2 0.328 95.8-131.6-108.3 2.5 43.7 103.5 38.3 101 104 A D S S+ 0 0 128 -5,-0.4 -4,-0.2 -6,-0.2 -3,-0.1 0.811 78.0 90.0 49.1 34.2 40.4 104.4 40.1 102 105 A I S S- 0 0 102 -6,-0.8 -2,-0.3 0, 0.0 -1,-0.2 -0.980 87.5 -84.9-151.6 156.2 39.9 107.2 37.6 103 106 A P - 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