==== Secondary Structure Definition by the program DSSP, updated CMBI version by ElmK / April 1,2000 ==== DATE=23-AUG-2013 . REFERENCE W. KABSCH AND C.SANDER, BIOPOLYMERS 22 (1983) 2577-2637 . HEADER CYTOKINE 19-MAR-13 4JPH . COMPND 2 MOLECULE: GREMLIN-2; . SOURCE 2 ORGANISM_SCIENTIFIC: MUS MUSCULUS; . AUTHOR X.DENG,K.T.NOLAN,C.KATTAMURI,T.B.THOMPSON . 438 4 16 16 0 TOTAL NUMBER OF RESIDUES, NUMBER OF CHAINS, NUMBER OF SS-BRIDGES(TOTAL,INTRACHAIN,INTERCHAIN) . 24942.0 ACCESSIBLE SURFACE OF PROTEIN (ANGSTROM**2) . 278 63.5 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(J) , SAME NUMBER PER 100 RESIDUES . 8 1.8 TOTAL NUMBER OF HYDROGEN BONDS IN PARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 159 36.3 TOTAL NUMBER OF HYDROGEN BONDS IN ANTIPARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-5), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-4), SAME NUMBER PER 100 RESIDUES . 1 0.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-3), SAME NUMBER PER 100 RESIDUES . 2 0.5 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-2), SAME NUMBER PER 100 RESIDUES . 1 0.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-1), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+0), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+1), SAME NUMBER PER 100 RESIDUES . 48 11.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+2), SAME NUMBER PER 100 RESIDUES . 25 5.7 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+3), SAME NUMBER PER 100 RESIDUES . 24 5.5 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+4), SAME NUMBER PER 100 RESIDUES . 2 0.5 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+5), SAME NUMBER PER 100 RESIDUES . 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 *** HISTOGRAMS OF *** . 0 0 0 2 1 0 0 1 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 RESIDUES PER ALPHA HELIX . 4 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PARALLEL BRIDGES PER LADDER . 0 0 0 2 2 0 0 0 2 0 4 0 0 3 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 ANTIPARALLEL BRIDGES PER LADDER . 4 0 0 0 0 0 2 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 LADDERS PER SHEET . # RESIDUE AA STRUCTURE BP1 BP2 ACC N-H-->O O-->H-N N-H-->O O-->H-N TCO KAPPA ALPHA PHI PSI X-CA Y-CA Z-CA 1 46 A H 0 0 230 0, 0.0 2,-0.2 0, 0.0 3,-0.1 0.000 360.0 360.0 360.0 -43.5 -20.8 26.2 44.7 2 47 A H > + 0 0 95 1,-0.2 3,-1.4 2,-0.1 4,-0.1 -0.334 360.0 148.5-102.3 49.3 -23.1 24.0 46.9 3 48 A Q T >> + 0 0 127 1,-0.3 3,-1.5 -2,-0.2 4,-0.9 0.449 48.2 94.8 -60.3 -5.6 -25.9 23.6 44.3 4 49 A I H 3> + 0 0 24 1,-0.3 4,-2.6 2,-0.2 -1,-0.3 0.573 65.5 77.3 -63.9 -15.4 -26.6 20.1 45.8 5 50 A K H <> S+ 0 0 133 -3,-1.4 4,-0.6 2,-0.2 -1,-0.3 0.845 96.9 45.4 -63.1 -35.1 -29.3 21.9 47.9 6 51 A E H X4 S+ 0 0 168 -3,-1.5 3,-0.5 2,-0.2 4,-0.3 0.896 111.3 51.5 -74.2 -42.2 -31.4 21.8 44.7 7 52 A V H >< S+ 0 0 36 -4,-0.9 3,-0.6 1,-0.2 -2,-0.2 0.927 118.0 39.2 -58.6 -44.0 -30.5 18.1 44.0 8 53 A L H >< S+ 0 0 0 -4,-2.6 3,-2.5 1,-0.2 6,-0.3 0.473 83.9 99.8 -86.9 -7.9 -31.5 17.2 47.6 9 54 A A T << S+ 0 0 66 -4,-0.6 -1,-0.2 -3,-0.5 -2,-0.1 0.735 79.0 61.4 -48.5 -31.8 -34.7 19.5 47.8 10 55 A S T < S+ 0 0 54 -3,-0.6 2,-0.6 -4,-0.3 -1,-0.3 0.636 100.6 57.2 -71.1 -20.8 -36.7 16.3 47.1 11 56 A S X - 0 0 1 -3,-2.5 3,-1.2 1,-0.2 4,-0.4 -0.834 65.0-176.1-118.6 94.2 -35.4 14.8 50.3 12 57 A Q G > S+ 0 0 147 -2,-0.6 3,-1.8 1,-0.2 4,-0.2 0.902 83.8 57.4 -49.5 -51.5 -36.4 17.1 53.3 13 58 A E G > S+ 0 0 52 1,-0.3 3,-0.5 2,-0.1 -1,-0.2 0.713 102.8 56.4 -54.5 -28.9 -34.5 14.8 55.8 14 59 A A G X S+ 0 0 0 -3,-1.2 3,-2.0 -6,-0.3 4,-0.4 0.444 75.7 91.0 -86.7 -6.3 -31.1 15.2 53.8 15 60 A L G X> + 0 0 46 -3,-1.8 3,-1.0 -4,-0.4 4,-0.5 0.693 69.2 80.1 -66.8 -11.2 -31.0 19.1 53.8 16 61 A V G <4 S+ 0 0 37 -3,-0.5 3,-0.3 1,-0.3 -1,-0.3 0.762 88.7 56.3 -58.1 -30.0 -28.9 18.6 57.1 17 62 A V G <4 S+ 0 0 0 -3,-2.0 -1,-0.3 1,-0.2 -2,-0.2 0.738 107.0 48.0 -74.2 -29.6 -26.1 17.9 54.5 18 63 A T T <4 S+ 0 0 21 -3,-1.0 -2,-0.2 -4,-0.4 -1,-0.2 0.440 88.1 112.0 -91.2 -9.0 -26.7 21.3 53.0 19 64 A E S >X S- 0 0 75 -4,-0.5 4,-2.8 -3,-0.3 3,-1.3 -0.446 75.9-125.2 -69.2 144.0 -26.7 23.2 56.4 20 65 A R H 3> S+ 0 0 180 1,-0.2 4,-2.9 2,-0.2 5,-0.2 0.861 108.6 65.6 -55.0 -39.5 -23.8 25.6 57.1 21 66 A K H 34 S+ 0 0 163 2,-0.2 -1,-0.2 1,-0.2 -4,-0.1 0.737 116.0 27.9 -58.3 -26.9 -23.0 23.7 60.3 22 67 A Y H X4 S+ 0 0 9 -3,-1.3 3,-1.4 -6,-0.3 4,-0.5 0.871 122.6 49.2 -95.7 -58.7 -22.1 20.6 58.3 23 68 A L H 3< S+ 0 0 16 -4,-2.8 -3,-0.2 1,-0.3 -2,-0.2 0.712 121.8 35.3 -53.7 -31.7 -20.9 22.2 55.0 24 69 A K T 3< S+ 0 0 119 -4,-2.9 -1,-0.3 -5,-0.2 -3,-0.2 0.358 102.2 73.8-106.9 3.0 -18.6 24.7 56.8 25 70 A S S < S+ 0 0 51 -3,-1.4 -2,-0.2 -5,-0.2 -1,-0.1 0.462 72.4 104.6 -93.0 -1.7 -17.5 22.4 59.7 26 71 A D - 0 0 52 -4,-0.5 2,-0.3 -3,-0.1 30,-0.2 -0.414 53.4-169.1 -75.1 151.8 -15.1 20.3 57.4 27 72 A W E -A 55 0A 74 28,-2.3 28,-1.9 -2,-0.1 2,-0.3 -0.992 22.5-177.2-148.6 153.7 -11.4 21.0 57.8 28 73 A a E -A 54 0A 26 -2,-0.3 26,-0.3 26,-0.2 2,-0.3 -0.848 17.5-157.0-154.5 108.9 -8.1 20.2 56.1 29 74 A K E -A 53 0A 114 24,-2.9 24,-1.4 -2,-0.3 2,-0.5 -0.664 8.5-150.2 -94.0 149.4 -4.8 21.4 57.6 30 75 A T E -A 52 0A 41 -2,-0.3 22,-0.2 22,-0.2 -2,-0.0 -0.973 24.5-179.8-113.4 122.3 -1.4 21.9 55.8 31 76 A Q E -A 51 0A 7 20,-2.6 20,-2.7 -2,-0.5 88,-0.1 -0.994 27.7-110.0-132.1 136.3 1.6 21.4 58.2 32 77 A P E -A 50 0A 16 0, 0.0 88,-1.4 0, 0.0 2,-0.3 -0.340 32.0-174.2 -67.7 147.4 5.4 21.6 57.6 33 78 A L E -Ab 49 120A 0 16,-2.5 16,-3.2 86,-0.2 2,-0.3 -0.997 22.7-126.5-139.9 134.7 7.5 18.5 57.6 34 79 A R E -A 48 0A 114 86,-2.9 2,-0.4 -2,-0.3 14,-0.2 -0.680 27.3-170.6 -85.5 134.4 11.3 18.2 57.4 35 80 A Q E -A 47 0A 6 12,-2.7 12,-2.9 -2,-0.3 2,-0.4 -0.998 15.1-141.4-125.6 135.1 12.6 15.9 54.6 36 81 A T E -A 46 0A 77 -2,-0.4 2,-0.5 10,-0.2 10,-0.2 -0.783 11.7-159.4 -93.4 130.4 16.2 14.7 54.1 37 82 A V E +A 45 0A 0 8,-3.4 8,-2.1 -2,-0.4 2,-0.3 -0.963 23.7 165.3-112.6 117.6 17.4 14.5 50.5 38 83 A S + 0 0 66 -2,-0.5 2,-0.3 6,-0.2 5,-0.1 -0.823 19.7 176.8-129.8 166.3 20.4 12.2 50.0 39 84 A E > - 0 0 58 3,-0.4 3,-2.3 -2,-0.3 2,-0.1 -0.859 44.9 -92.6-169.0 138.7 22.5 10.4 47.4 40 85 A E T 3 S+ 0 0 183 1,-0.3 3,-0.1 -2,-0.3 52,-0.0 -0.403 116.1 20.8 -61.7 122.9 25.6 8.3 47.6 41 86 A G T 3 S+ 0 0 32 1,-0.3 53,-1.8 -2,-0.1 54,-0.4 0.308 110.6 99.4 97.3 -2.0 28.6 10.5 47.0 42 87 A b S < S- 0 0 19 -3,-2.3 2,-0.5 50,-0.2 -3,-0.4 -0.839 78.1-113.0-115.4 156.0 26.6 13.7 48.0 43 88 A R - 0 0 148 48,-2.8 48,-0.4 -2,-0.3 -6,-0.0 -0.713 43.8-122.7 -83.5 123.1 26.2 15.9 51.1 44 89 A S - 0 0 64 -2,-0.5 2,-0.4 46,-0.1 -6,-0.2 -0.238 24.4-156.9 -65.8 153.0 22.6 15.5 52.3 45 90 A R E -A 37 0A 101 -8,-2.1 -8,-3.4 43,-0.0 2,-0.5 -0.980 10.2-138.7-132.7 147.0 20.3 18.4 52.7 46 91 A T E +A 36 0A 87 -2,-0.4 2,-0.3 -10,-0.2 -10,-0.2 -0.921 27.4 179.6-104.6 131.6 17.2 18.9 54.8 47 92 A I E -A 35 0A 27 -12,-2.9 -12,-2.7 -2,-0.5 2,-0.5 -0.775 28.7-111.9-122.7 169.0 14.2 20.7 53.2 48 93 A L E +A 34 0A 78 -2,-0.3 2,-0.2 -14,-0.2 -14,-0.2 -0.917 38.4 165.3-105.4 126.1 10.7 21.7 54.2 49 94 A N E -A 33 0A 0 -16,-3.2 -16,-2.5 -2,-0.5 2,-0.3 -0.635 28.7-121.0-119.8-177.0 7.8 20.0 52.6 50 95 A R E -A 32 0A 76 -18,-0.3 57,-0.3 -2,-0.2 2,-0.3 -0.958 15.0-155.9-129.9 150.9 4.1 20.0 53.8 51 96 A F E -A 31 0A 0 -20,-2.7 -20,-2.6 -2,-0.3 2,-0.3 -0.878 12.5-134.5-122.3 157.8 1.7 17.2 54.8 52 97 A c E +AC 30 172A 0 120,-3.1 120,-2.4 -2,-0.3 2,-0.3 -0.894 26.6 160.6-109.9 145.2 -2.2 17.1 54.8 53 98 A Y E +AC 29 171A 6 -24,-1.4 -24,-2.9 -2,-0.3 2,-0.3 -0.928 19.3 113.5-161.7 135.8 -4.6 15.9 57.5 54 99 A G E -AC 28 170A 0 116,-2.4 116,-1.9 -2,-0.3 2,-0.4 -0.968 56.9 -96.5-176.1-171.6 -8.3 16.6 58.1 55 100 A Q E -AC 27 169A 13 -28,-1.9 -28,-2.3 -2,-0.3 2,-0.2 -0.937 41.5-175.5-136.5 104.4 -12.0 15.5 58.2 56 101 A d E - C 0 168A 3 112,-0.8 112,-0.8 -2,-0.4 2,-0.3 -0.552 31.9 -88.8 -99.3 164.8 -13.9 16.3 55.0 57 102 A N E + C 0 167A 13 -2,-0.2 19,-1.8 110,-0.2 2,-0.3 -0.577 46.1 166.7 -81.3 135.5 -17.5 15.8 54.2 58 103 A S E +DC 75 166A 1 108,-1.9 108,-2.1 -2,-0.3 2,-0.3 -0.988 4.0 165.8-145.9 139.6 -18.8 12.5 52.7 59 104 A F E -DC 74 165A 8 15,-2.1 15,-2.0 -2,-0.3 2,-0.3 -0.976 24.6-154.1-149.9 163.3 -22.3 11.3 52.4 60 105 A Y E -DC 73 164A 35 104,-2.3 104,-2.7 -2,-0.3 13,-0.2 -0.949 17.5-178.4-145.1 114.4 -24.4 8.6 50.7 61 106 A I E -D 72 0A 0 11,-3.5 11,-2.7 -2,-0.3 2,-0.2 -0.941 22.3-140.1-122.7 112.1 -28.1 9.3 50.0 62 107 A P E -D 71 0A 7 0, 0.0 9,-0.2 0, 0.0 2,-0.2 -0.485 30.8-146.7 -67.6 128.1 -30.5 6.8 48.4 63 108 A R - 0 0 52 7,-1.9 7,-0.3 -2,-0.2 84,-0.1 -0.667 27.6 -74.9 -98.1 161.4 -32.7 8.8 46.0 64 109 A H - 0 0 38 -2,-0.2 2,-1.1 1,-0.1 6,-0.2 -0.158 51.2-121.4 -48.7 135.3 -36.3 8.5 44.9 65 110 A V B +e 149 0A 1 83,-2.8 85,-3.4 1,-0.2 4,-0.1 -0.817 56.0 143.0 -93.4 100.5 -36.5 5.5 42.5 66 111 A K S S- 0 0 131 -2,-1.1 86,-0.2 2,-0.6 -1,-0.2 0.829 80.5 -12.6-102.6 -54.2 -37.9 7.0 39.3 67 112 A K S S- 0 0 103 1,-0.6 2,-0.2 84,-0.1 -2,-0.1 0.391 132.4 -0.3-134.5 1.9 -36.2 5.3 36.3 68 113 A E S S- 0 0 70 2,-0.0 -1,-0.6 83,-0.0 -2,-0.6 -0.815 86.0 -87.3 176.0 148.8 -33.2 3.5 37.8 69 114 A E + 0 0 97 -2,-0.2 2,-0.3 -4,-0.1 -4,-0.1 -0.627 59.7 162.5 -75.3 115.2 -31.8 3.0 41.4 70 115 A D - 0 0 121 -2,-0.6 -7,-1.9 -7,-0.3 2,-0.3 -0.831 21.2-162.2-128.7 163.8 -29.4 6.0 41.9 71 116 A S E -D 62 0A 30 -2,-0.3 2,-0.6 -9,-0.2 -2,-0.0 -0.985 23.0-122.4-149.4 141.6 -28.0 7.6 45.0 72 117 A F E +D 61 0A 46 -11,-2.7 -11,-3.5 -2,-0.3 2,-0.3 -0.799 40.1 174.5 -87.0 124.5 -26.4 10.9 45.6 73 118 A Q E -D 60 0A 88 -2,-0.6 2,-0.3 -13,-0.2 -13,-0.2 -0.956 15.9-174.1-129.2 147.5 -22.8 10.3 46.9 74 119 A S E -D 59 0A 31 -15,-2.0 -15,-2.1 -2,-0.3 2,-0.2 -0.977 1.3-175.5-142.6 133.6 -20.0 12.8 47.7 75 120 A C E +D 58 0A 29 -2,-0.3 -17,-0.2 -17,-0.2 2,-0.2 -0.626 10.3 172.3-130.8 74.1 -16.5 11.7 48.8 76 121 A A - 0 0 13 -19,-1.8 2,-0.3 -2,-0.2 38,-0.2 -0.526 7.8-174.8 -84.9 152.3 -14.2 14.6 49.7 77 122 A F E -H 113 0B 30 36,-1.6 36,-2.3 -2,-0.2 2,-0.9 -0.983 33.3-107.8-145.8 148.3 -10.7 13.9 51.2 78 123 A a E +H 112 0B 1 -2,-0.3 34,-0.2 34,-0.2 -24,-0.2 -0.695 63.4 136.2 -81.0 102.8 -7.9 16.1 52.6 79 124 A K E -H 111 0B 74 32,-1.8 32,-2.3 -2,-0.9 2,-0.1 -0.901 61.3 -63.0-143.6 167.8 -5.2 16.0 50.0 80 125 A P E -H 110 0B 13 0, 0.0 30,-0.3 0, 0.0 3,-0.2 -0.389 32.9-159.6 -61.6 130.0 -2.7 18.3 48.1 81 126 A Q E S+ 0 0 75 28,-3.8 2,-0.3 1,-0.4 29,-0.2 0.858 86.9 13.3 -71.2 -39.2 -4.6 20.9 46.1 82 127 A R E -H 109 0B 129 27,-0.8 26,-1.8 -3,-0.0 27,-0.7 -0.988 67.7-167.7-142.8 123.6 -1.4 21.4 44.0 83 128 A V E -H 107 0B 71 -2,-0.3 2,-0.3 -3,-0.2 24,-0.2 -0.984 18.7-169.3-112.4 119.0 1.7 19.0 44.1 84 129 A T E -H 106 0B 68 22,-2.5 22,-3.3 -2,-0.5 2,-0.6 -0.820 25.1-119.0-113.4 155.6 4.8 20.6 42.3 85 130 A S E -H 105 0B 66 -2,-0.3 2,-0.4 20,-0.2 20,-0.2 -0.843 35.3-167.7 -93.3 120.6 8.1 19.2 41.2 86 131 A V E -H 104 0B 42 18,-3.2 18,-3.0 -2,-0.6 2,-0.3 -0.942 17.8-128.5-114.5 137.3 10.9 21.1 43.0 87 132 A I E -H 103 0B 83 -2,-0.4 2,-0.5 16,-0.2 16,-0.2 -0.647 19.2-169.1 -81.1 130.9 14.7 21.0 42.3 88 133 A V E -H 102 0B 5 14,-2.8 14,-2.9 -2,-0.3 2,-0.3 -0.974 14.8-145.7-123.3 112.3 17.0 20.2 45.2 89 134 A E E -H 101 0B 107 -2,-0.5 2,-0.4 12,-0.2 12,-0.2 -0.600 14.9-165.4 -78.6 137.1 20.7 20.9 44.4 90 135 A L E -H 100 0B 5 10,-2.9 10,-2.0 -2,-0.3 2,-0.5 -0.984 18.6-137.3-125.8 140.2 23.3 18.6 46.0 91 136 A E E -H 99 0B 110 -48,-0.4 -48,-2.8 -2,-0.4 8,-0.3 -0.827 22.8-170.5 -91.6 130.5 27.0 19.1 46.2 92 137 A b > + 0 0 0 6,-2.5 3,-2.4 -2,-0.5 -50,-0.2 -0.667 17.3 170.8-129.4 74.3 28.6 15.6 45.4 93 138 A P T 3 S+ 0 0 99 0, 0.0 -51,-0.1 0, 0.0 -1,-0.1 0.804 83.5 53.9 -54.1 -32.4 32.4 15.6 46.2 94 139 A G T 3 S+ 0 0 60 -53,-1.8 2,-0.3 4,-0.1 -52,-0.1 0.335 103.8 69.5 -82.2 2.4 32.5 11.8 45.6 95 140 A L S < S- 0 0 50 -3,-2.4 -3,-0.1 -54,-0.4 -54,-0.1 -0.783 82.0-101.3-123.6 168.4 30.9 12.0 42.1 96 141 A D S S+ 0 0 92 -2,-0.3 -2,-0.0 2,-0.1 0, 0.0 -0.993 114.7 29.0-130.2 119.8 31.6 13.1 38.6 97 142 A P S S- 0 0 2 0, 0.0 2,-1.2 0, 0.0 -1,-0.1 0.684 104.8-125.4 -65.2 159.9 30.4 15.9 37.8 98 143 A P S S+ 0 0 49 0, 0.0 -6,-2.5 0, 0.0 2,-0.3 0.033 81.8 56.6 -78.2 40.3 30.6 16.9 41.5 99 144 A F E -H 91 0B 32 -2,-1.2 2,-0.3 -8,-0.3 -8,-0.2 -0.984 58.3-163.6-160.1 156.8 26.9 17.8 41.8 100 145 A R E -H 90 0B 27 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