==== Secondary Structure Definition by the program DSSP, updated CMBI version by ElmK / April 1,2000 ==== DATE=6-DEC-2009 . REFERENCE W. KABSCH AND C.SANDER, BIOPOLYMERS 22 (1983) 2577-2637 . HEADER STRUCTURAL PROTEIN 17-APR-02 1LI1 . COMPND 2 MOLECULE: COLLAGEN ALPHA 1(IV); . SOURCE 2 ORGANISM_SCIENTIFIC: HOMO SAPIENS; . AUTHOR M.E.THAN,S.HENRICH,R.HUBER,A.RIES,K.MANN,K.KUHN,R.TIMPL, . 1359 6 36 36 0 TOTAL NUMBER OF RESIDUES, NUMBER OF CHAINS, NUMBER OF SS-BRIDGES(TOTAL,INTRACHAIN,INTERCHAIN) . 39554.0 ACCESSIBLE SURFACE OF PROTEIN (ANGSTROM**2) . 824 60.6 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(J) , SAME NUMBER PER 100 RESIDUES . 4 0.3 TOTAL NUMBER OF HYDROGEN BONDS IN PARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 522 38.4 TOTAL NUMBER OF HYDROGEN BONDS IN ANTIPARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 2 0.1 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-5), SAME NUMBER PER 100 RESIDUES . 16 1.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-4), SAME NUMBER PER 100 RESIDUES . 6 0.4 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-3), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-2), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-1), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+0), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+1), SAME NUMBER PER 100 RESIDUES . 96 7.1 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+2), SAME NUMBER PER 100 RESIDUES . 146 10.7 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+3), SAME NUMBER PER 100 RESIDUES . 13 1.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+4), SAME NUMBER PER 100 RESIDUES . 2 0.1 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+5), SAME NUMBER PER 100 RESIDUES . 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 *** HISTOGRAMS OF *** . 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 RESIDUES PER ALPHA HELIX . 2 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PARALLEL BRIDGES PER LADDER . 12 24 11 31 13 11 6 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 ANTIPARALLEL BRIDGES PER LADDER . 0 0 12 0 12 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 LADDERS PER SHEET . # RESIDUE AA STRUCTURE BP1 BP2 ACC N-H-->O O-->H-N N-H-->O O-->H-N TCO KAPPA ALPHA PHI PSI X-CA Y-CA Z-CA 1 2 A V 0 0 154 0, 0.0 2,-0.6 0, 0.0 455,-0.1 0.000 360.0 360.0 360.0 38.7 89.9 53.3 91.3 2 3 A D - 0 0 68 1,-0.1 112,-0.1 453,-0.1 109,-0.0 -0.772 360.0-174.3 -88.5 116.8 86.4 53.6 89.8 3 4 A H - 0 0 24 -2,-0.6 340,-3.0 110,-0.4 451,-0.3 0.523 53.9 -93.2 -89.1 -8.4 86.2 51.5 86.6 4 5 A G + 0 0 1 108,-0.3 2,-0.2 1,-0.2 450,-0.1 0.798 56.7 173.5 101.8 35.8 82.7 52.6 85.6 5 6 A F E -A 112 0A 22 107,-0.9 107,-2.7 338,-0.1 2,-0.4 -0.554 15.1-157.6 -76.2 139.9 80.3 50.1 87.1 6 7 A L E -A 111 0A 44 105,-0.3 2,-0.4 -2,-0.2 105,-0.2 -0.919 16.6-176.3-122.5 146.6 76.6 51.0 86.6 7 8 A V E -A 110 0A 0 103,-2.9 103,-2.7 -2,-0.4 2,-0.4 -0.993 12.8-152.8-139.6 131.7 73.4 50.1 88.4 8 9 A T E -A 109 0A 1 39,-1.2 2,-0.4 -2,-0.4 101,-0.2 -0.874 10.3-165.4-106.6 137.6 70.0 51.2 87.3 9 10 A R E -A 108 0A 42 99,-2.5 99,-2.4 -2,-0.4 2,-0.5 -0.934 6.1-152.6-123.5 145.1 67.2 51.5 89.9 10 11 A H E -A 107 0A 11 -2,-0.4 155,-0.3 97,-0.2 97,-0.2 -0.970 6.9-161.2-120.2 129.2 63.4 51.8 89.2 11 12 A S - 0 0 20 95,-2.6 96,-0.1 -2,-0.5 3,-0.1 0.793 22.3-144.2 -78.2 -26.5 61.1 53.6 91.6 12 13 A Q S S+ 0 0 0 94,-0.7 2,-0.3 1,-0.3 93,-0.1 0.779 76.3 78.8 64.5 24.1 58.0 52.0 90.2 13 14 A T S S- 0 0 44 2,-0.4 93,-1.5 93,-0.2 -1,-0.3 -0.874 100.2 -97.5-145.2 177.8 56.3 55.4 91.0 14 15 A I S S+ 0 0 99 -2,-0.3 2,-0.3 91,-0.2 89,-0.1 0.499 98.6 87.4 -82.0 -1.0 56.3 58.8 89.3 15 16 A D S S- 0 0 118 91,-0.1 91,-0.4 1,-0.1 -2,-0.4 -0.690 90.6-103.4 -94.3 152.2 59.0 60.0 91.7 16 17 A D - 0 0 72 -2,-0.3 2,-0.1 89,-0.1 92,-0.1 -0.534 35.5-125.2 -71.5 135.4 62.7 59.6 90.8 17 18 A P - 0 0 13 0, 0.0 2,-0.2 0, 0.0 -8,-0.1 -0.483 26.1-116.6 -77.3 153.5 64.3 56.7 92.8 18 19 A Q - 0 0 160 -2,-0.1 90,-0.1 1,-0.1 3,-0.1 -0.632 28.6-106.7 -86.9 150.1 67.4 57.7 94.8 19 20 A a - 0 0 24 -2,-0.2 3,-0.1 1,-0.1 6,-0.1 -0.537 48.1-102.9 -70.4 143.2 70.7 56.1 94.0 20 21 A P > - 0 0 6 0, 0.0 3,-2.1 0, 0.0 -1,-0.1 -0.266 48.2 -76.7 -67.7 155.9 71.5 53.6 96.7 21 22 A S T 3 S+ 0 0 101 1,-0.3 3,-0.1 115,-0.3 -3,-0.0 -0.211 121.0 29.5 -53.3 137.6 74.1 54.4 99.4 22 23 A G T 3 S+ 0 0 58 1,-0.3 2,-0.3 -3,-0.1 -1,-0.3 0.350 105.6 95.9 93.2 -6.5 77.6 54.1 98.1 23 24 A T < - 0 0 24 -3,-2.1 2,-0.5 89,-0.1 -1,-0.3 -0.808 66.2-131.4-118.1 161.9 76.6 55.0 94.5 24 25 A K E -B 111 0A 85 87,-2.8 87,-3.2 -2,-0.3 2,-0.2 -0.920 21.4-122.1-114.9 130.5 76.6 58.2 92.5 25 26 A I E +B 110 0A 73 -2,-0.5 85,-0.2 85,-0.2 3,-0.1 -0.452 29.7 171.9 -70.1 135.0 73.7 59.5 90.5 26 27 A L E - 0 0 26 83,-2.8 2,-0.3 1,-0.5 333,-0.2 0.749 62.4 -39.4-105.5 -52.0 74.4 60.0 86.8 27 28 A Y E -B 109 0A 32 82,-1.3 82,-3.2 331,-0.1 -1,-0.5 -0.977 50.3-126.6-166.6 171.0 70.9 60.8 85.4 28 29 A H E +B 108 0A 59 -2,-0.3 2,-0.3 80,-0.2 80,-0.2 -0.735 31.5 144.7-121.6 172.1 67.2 59.8 85.7 29 30 A G E -B 107 0A 0 78,-1.9 78,-2.3 -2,-0.2 2,-0.3 -0.921 47.4 -53.6-175.7-159.1 64.6 58.8 83.2 30 31 A Y E -BC 106 53A 33 23,-3.3 23,-2.1 -2,-0.3 2,-1.0 -0.830 52.8-108.2-106.8 143.5 61.6 56.6 82.5 31 32 A S E - C 0 52A 0 74,-2.4 51,-2.0 -2,-0.3 2,-0.5 -0.565 34.5-170.5 -76.1 96.7 61.5 52.9 83.0 32 33 A L E +D 81 0B 4 19,-2.1 49,-0.3 -2,-1.0 12,-0.1 -0.742 11.7 175.3 -89.3 128.0 61.5 51.1 79.6 33 34 A L E - 0 0 0 47,-2.7 11,-2.3 -2,-0.5 2,-0.3 0.869 59.4 -43.3 -95.7 -52.9 60.8 47.3 79.9 34 35 A Y E -DE 80 43B 1 46,-1.4 46,-3.0 9,-0.3 -1,-0.4 -0.982 44.7-132.6-168.4 170.6 60.6 46.4 76.1 35 36 A V E -DE 79 42B 6 7,-2.3 7,-2.6 -2,-0.3 2,-0.4 -0.885 9.5-148.8-129.9 161.3 59.3 47.2 72.7 36 37 A Q E +DE 78 41B 0 42,-2.4 42,-2.5 -2,-0.3 2,-0.4 -0.970 19.1 175.0-135.9 116.3 57.6 45.2 70.0 37 38 A G E > S- E 0 40B 0 3,-2.2 3,-1.9 -2,-0.4 40,-0.1 -0.990 72.7 -2.9-125.7 134.1 58.1 46.1 66.4 38 39 A N T 3 S- 0 0 6 -2,-0.4 -1,-0.2 1,-0.3 793,-0.1 0.894 133.1 -58.2 51.2 40.3 56.8 44.1 63.4 39 40 A E T 3 S+ 0 0 13 37,-0.2 2,-0.5 1,-0.2 -1,-0.3 0.512 113.1 116.8 69.9 10.6 55.4 41.7 66.1 40 41 A R E < -E 37 0B 48 -3,-1.9 -3,-2.2 111,-0.1 2,-0.3 -0.927 56.5-139.0-112.8 133.1 58.8 41.0 67.6 41 42 A A E -E 36 0B 3 -2,-0.5 2,-0.4 -5,-0.2 -5,-0.2 -0.623 18.5-179.8 -88.2 142.6 59.7 41.8 71.2 42 43 A H E -E 35 0B 26 -7,-2.6 -7,-2.3 -2,-0.3 2,-0.2 -0.946 14.3-163.4-140.6 117.5 63.1 43.2 72.2 43 44 A G E -E 34 0B 20 -2,-0.4 2,-0.4 -9,-0.3 -9,-0.3 -0.502 16.3-136.4-101.7 171.4 63.7 44.0 75.9 44 45 A Q - 0 0 6 -11,-2.3 2,-0.2 -2,-0.2 340,-0.1 -0.995 33.1-112.7-124.8 128.2 66.1 46.0 78.0 45 46 A D > - 0 0 34 -2,-0.4 3,-2.4 1,-0.2 6,-0.5 -0.435 22.4-136.7 -63.9 126.4 67.5 44.4 81.2 46 47 A L T 3 S+ 0 0 1 1,-0.3 -1,-0.2 -2,-0.2 109,-0.1 0.629 101.5 61.4 -61.2 -14.7 66.1 46.5 84.0 47 48 A G T 3 S+ 0 0 5 -40,-0.1 -39,-1.2 109,-0.1 -1,-0.3 0.546 97.2 76.3 -88.6 -5.2 69.5 46.5 85.8 48 49 A T S X S- 0 0 47 -3,-2.4 3,-2.1 -41,-0.2 4,-0.1 -0.700 88.9-116.3-108.0 159.5 71.1 48.3 82.8 49 50 A A G > S+ 0 0 5 1,-0.3 3,-1.9 -2,-0.2 -1,-0.1 0.675 107.4 77.1 -63.8 -16.8 70.9 52.0 81.7 50 51 A G G 3 S+ 0 0 1 1,-0.3 -1,-0.3 -5,-0.2 -4,-0.1 0.709 88.9 56.9 -67.7 -18.5 69.2 50.8 78.5 51 52 A S G < S+ 0 0 0 -3,-2.1 -19,-2.1 -6,-0.5 2,-0.5 0.348 96.9 73.2 -91.8 6.5 66.0 50.4 80.6 52 53 A b E < -C 31 0A 3 -3,-1.9 2,-0.3 -21,-0.2 -21,-0.2 -0.829 65.0-179.1-125.6 93.3 66.2 54.1 81.6 53 54 A L E -C 30 0A 7 -23,-2.1 -23,-3.3 -2,-0.5 54,-0.1 -0.720 29.5-139.0 -95.8 142.3 65.3 56.5 78.8 54 55 A R S S+ 0 0 64 294,-0.4 2,-0.4 -2,-0.3 -1,-0.1 0.831 88.8 44.6 -66.1 -30.8 65.4 60.3 79.1 55 56 A K S S- 0 0 44 -25,-0.1 2,-0.3 5,-0.1 -1,-0.1 -0.933 71.8-146.3-122.3 140.6 62.2 60.7 77.2 56 57 A F + 0 0 0 -2,-0.4 2,-0.3 369,-0.1 3,-0.0 -0.711 26.3 154.1 -97.5 152.1 58.9 58.8 77.3 57 58 A S - 0 0 1 -2,-0.3 371,-0.1 372,-0.1 372,-0.1 -0.892 49.7-128.3-167.2 146.0 56.6 58.1 74.3 58 59 A T S S+ 0 0 3 370,-0.4 120,-0.1 372,-0.3 117,-0.1 0.745 116.4 35.8 -69.4 -18.6 54.1 55.4 73.5 59 60 A M + 0 0 2 118,-0.1 -1,-0.3 1,-0.1 362,-0.1 -0.751 68.9 164.8-134.5 85.0 56.1 55.1 70.3 60 61 A P + 0 0 0 0, 0.0 361,-2.2 0, 0.0 2,-0.3 0.455 61.7 48.4 -81.5 2.7 59.8 55.6 71.1 61 62 A F E -I 420 0C 2 359,-0.3 12,-0.5 385,-0.0 2,-0.3 -0.858 58.1-159.0-138.8 169.1 61.1 54.2 67.8 62 63 A L E -IJ 419 72C 5 357,-2.2 357,-2.0 -2,-0.3 2,-0.3 -0.877 17.8-133.6-136.8 167.6 60.6 54.2 64.1 63 64 A F E -IJ 418 71C 9 8,-2.2 8,-3.0 -2,-0.3 2,-0.3 -0.960 13.3-163.2-131.4 153.0 61.6 51.7 61.4 64 65 A c E -IJ 417 70C 0 353,-2.3 353,-2.1 -2,-0.3 6,-0.2 -0.916 5.2-151.1-131.1 156.6 63.2 52.0 58.0 65 66 A N > - 0 0 13 4,-2.1 3,-1.7 -2,-0.3 4,-0.4 -0.687 37.7 -94.6-123.8 175.9 63.5 49.8 54.9 66 67 A I T 3 S+ 0 0 26 349,-0.3 332,-0.2 346,-0.3 331,-0.1 0.603 117.3 69.0 -67.3 -9.7 66.0 49.2 52.1 67 68 A N T 3 S- 0 0 93 2,-0.2 -1,-0.3 346,-0.2 372,-0.3 0.260 116.1-110.3 -92.0 13.5 64.0 51.7 49.9 68 69 A N S < S+ 0 0 81 -3,-1.7 371,-0.5 1,-0.2 2,-0.4 0.905 77.6 129.0 58.4 44.7 65.2 54.6 52.1 69 70 A V E - K 0 438C 14 -4,-0.4 -4,-2.1 369,-0.2 2,-0.4 -0.990 38.3-167.0-126.6 133.8 61.7 55.1 53.6 70 71 A c E -JK 64 437C 0 367,-2.3 367,-3.3 -2,-0.4 2,-0.4 -0.954 3.8-157.8-121.8 146.4 61.2 55.3 57.4 71 72 A N E -JK 63 436C 12 -8,-3.0 -8,-2.2 -2,-0.4 2,-0.5 -0.989 2.4-164.4-126.7 131.0 57.8 55.2 59.1 72 73 A F E S-JK 62 435C 24 363,-2.9 363,-1.9 -2,-0.4 -10,-0.2 -0.974 76.8 -16.8-118.3 126.8 57.1 56.5 62.6 73 74 A A S S+ 0 0 12 -12,-0.5 -1,-0.2 -2,-0.5 -11,-0.1 0.835 101.7 122.8 49.8 38.0 54.0 55.5 64.4 74 75 A S + 0 0 3 361,-0.1 2,-0.1 -3,-0.1 -1,-0.1 -0.059 65.4 28.2-121.4 31.9 52.6 54.4 61.1 75 76 A R S S- 0 0 8 2,-0.1 787,-0.1 701,-0.0 103,-0.0 -0.369 104.1 -66.9-149.6-122.7 51.8 50.7 61.7 76 77 A N + 0 0 3 -2,-0.1 -37,-0.2 101,-0.1 2,-0.2 -0.465 69.3 144.8-146.8 65.6 50.8 48.7 64.8 77 78 A D - 0 0 10 -39,-0.1 100,-2.4 -40,-0.1 2,-0.3 -0.564 30.5-140.7-109.7 171.9 53.7 48.6 67.2 78 79 A Y E -DF 36 176B 0 -42,-2.5 -42,-2.4 98,-0.2 2,-0.3 -0.887 10.7-165.2-131.0 159.0 54.1 48.6 71.0 79 80 A S E -DF 35 175B 0 96,-2.1 96,-3.1 -2,-0.3 2,-0.4 -0.984 14.0-156.6-138.8 143.7 56.2 49.9 73.9 80 81 A Y E -DF 34 174B 0 -46,-3.0 -47,-2.7 -2,-0.3 -46,-1.4 -0.986 11.6-177.1-126.4 134.1 56.1 48.5 77.5 81 82 A W E -DF 32 173B 0 92,-1.8 92,-2.8 -2,-0.4 -49,-0.2 -0.951 35.1-103.1-131.7 147.3 57.1 50.5 80.5 82 83 A L E - F 0 172B 0 -51,-2.0 23,-2.5 -2,-0.3 90,-0.3 -0.437 50.9-144.4 -63.1 144.7 57.3 49.6 84.2 83 84 A S B -N 104 0D 0 88,-2.7 21,-0.2 21,-0.2 20,-0.1 -0.581 5.5-121.7-115.3 176.5 54.3 51.2 85.9 84 85 A T - 0 0 8 19,-1.3 -71,-0.2 -2,-0.2 -1,-0.1 -0.358 41.4 -92.0-103.4-178.5 53.3 52.8 89.1 85 86 A P S S+ 0 0 85 0, 0.0 18,-0.1 0, 0.0 -72,-0.1 0.502 77.7 134.8 -74.7 -3.2 50.6 52.0 91.7 86 87 A E - 0 0 85 16,-0.4 -2,-0.1 1,-0.1 2,-0.0 -0.219 59.6-118.4 -50.5 124.1 48.1 54.2 89.8 87 88 A P - 0 0 87 0, 0.0 -1,-0.1 0, 0.0 3,-0.1 -0.320 25.8-102.5 -69.5 148.6 44.8 52.3 89.5 88 89 A M - 0 0 63 1,-0.1 94,-0.0 2,-0.1 5,-0.0 -0.441 51.8 -93.6 -66.0 137.9 43.4 51.3 86.1 89 90 A P > - 0 0 44 0, 0.0 3,-1.6 0, 0.0 -1,-0.1 -0.190 31.9-116.2 -52.7 146.6 40.5 53.7 85.2 90 91 A M T 3 S+ 0 0 183 1,-0.3 -2,-0.1 2,-0.1 0, 0.0 0.836 116.8 58.5 -55.0 -35.3 37.1 52.4 86.2 91 92 A S T 3 S- 0 0 66 1,-0.1 -1,-0.3 2,-0.1 1027,-0.0 0.763 97.0-144.1 -65.9 -24.9 36.1 52.3 82.5 92 93 A M < + 0 0 64 -3,-1.6 -2,-0.1 1,-0.3 -1,-0.1 0.641 54.6 135.3 67.5 14.1 39.0 49.9 82.0 93 94 A A - 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