==== Secondary Structure Definition by the program DSSP, updated CMBI version by ElmK / April 1,2000 ==== DATE=6-JAN-2010 . REFERENCE W. KABSCH AND C.SANDER, BIOPOLYMERS 22 (1983) 2577-2637 . HEADER TRANSFERASE 06-MAY-07 2YZK . COMPND 2 MOLECULE: OROTATE PHOSPHORIBOSYLTRANSFERASE; . SOURCE 2 ORGANISM_SCIENTIFIC: AEROPYRUM PERNIX; . AUTHOR M.KANAGAWA,S.BABA,S.KURAMITSU,S.YOKOYAMA,G.KAWAI,G.SAMPEI, . 687 6 0 0 0 TOTAL NUMBER OF RESIDUES, NUMBER OF CHAINS, NUMBER OF SS-BRIDGES(TOTAL,INTRACHAIN,INTERCHAIN) . 27135.0 ACCESSIBLE SURFACE OF PROTEIN (ANGSTROM**2) . 515 75.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(J) , SAME NUMBER PER 100 RESIDUES . 115 16.7 TOTAL NUMBER OF HYDROGEN BONDS IN PARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 39 5.7 TOTAL NUMBER OF HYDROGEN BONDS IN ANTIPARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-5), SAME NUMBER PER 100 RESIDUES . 4 0.6 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-4), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-3), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-2), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-1), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+0), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+1), SAME NUMBER PER 100 RESIDUES . 36 5.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+2), SAME NUMBER PER 100 RESIDUES . 65 9.5 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+3), SAME NUMBER PER 100 RESIDUES . 205 29.8 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+4), SAME NUMBER PER 100 RESIDUES . 24 3.5 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+5), SAME NUMBER PER 100 RESIDUES . 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 *** HISTOGRAMS OF *** . 0 0 0 0 0 2 7 3 0 0 4 4 0 0 0 0 0 0 0 2 2 0 0 0 0 0 0 0 0 0 RESIDUES PER ALPHA HELIX . 0 0 0 8 0 4 1 3 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PARALLEL BRIDGES PER LADDER . 4 1 0 0 4 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 ANTIPARALLEL BRIDGES PER LADDER . 3 1 0 0 4 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 LADDERS PER SHEET . # RESIDUE AA STRUCTURE BP1 BP2 ACC N-H-->O O-->H-N N-H-->O O-->H-N TCO KAPPA ALPHA PHI PSI X-CA Y-CA Z-CA 1 1 A X >> 0 0 90 0, 0.0 4,-0.9 0, 0.0 3,-0.6 0.000 360.0 360.0 360.0 92.1 38.7 5.3 4.3 2 2 A L H 3> + 0 0 3 1,-0.2 4,-3.3 2,-0.2 5,-0.3 0.902 360.0 75.9 -56.5 -45.8 41.0 4.6 7.3 3 3 A A H 3> S+ 0 0 2 1,-0.2 4,-2.3 2,-0.2 -1,-0.2 0.813 97.0 45.5 -34.1 -53.5 38.8 1.9 8.7 4 4 A K H <> S+ 0 0 138 -3,-0.6 4,-2.2 1,-0.2 -1,-0.2 0.949 116.8 43.7 -60.7 -51.1 39.9 -0.6 6.1 5 5 A V H X S+ 0 0 23 -4,-0.9 4,-2.0 -3,-0.3 6,-0.3 0.871 112.4 54.3 -62.3 -39.1 43.6 0.2 6.4 6 6 A L H X>S+ 0 0 0 -4,-3.3 5,-2.6 2,-0.2 6,-0.8 0.920 109.4 46.5 -62.3 -45.0 43.3 0.3 10.2 7 7 A K H ><5S+ 0 0 126 -4,-2.3 3,-1.0 -5,-0.3 -2,-0.2 0.922 110.4 54.3 -63.5 -44.0 41.8 -3.2 10.4 8 8 A K H 3<5S+ 0 0 155 -4,-2.2 -1,-0.2 1,-0.3 -2,-0.2 0.878 113.9 40.2 -56.8 -43.8 44.4 -4.6 8.0 9 9 A R H 3<5S- 0 0 40 -4,-2.0 -1,-0.3 -5,-0.1 -2,-0.2 0.413 112.5-114.1 -88.5 1.0 47.3 -3.3 10.1 10 10 A G T <<5S+ 0 0 24 -3,-1.0 -3,-0.2 -4,-0.5 -4,-0.1 0.749 73.3 138.3 72.4 21.0 45.7 -4.2 13.4 11 11 A A S - 0 0 26 4,-2.7 3,-2.6 -2,-0.4 345,-0.1 -0.578 28.9-108.5 -96.7 162.1 41.7 2.8 29.2 20 20 A S T 3 S+ 0 0 39 1,-0.3 347,-0.2 -2,-0.2 356,-0.1 0.605 119.2 67.0 -62.6 -10.4 43.5 5.6 31.1 21 21 A S T 3 S- 0 0 41 2,-0.2 -1,-0.3 345,-0.1 3,-0.1 0.673 117.4-115.2 -81.5 -18.7 40.0 7.2 31.3 22 22 A G S < S+ 0 0 1 -3,-2.6 -2,-0.1 1,-0.4 2,-0.1 0.194 74.5 129.5 102.7 -13.8 39.0 4.4 33.5 23 23 A R - 0 0 27 -5,-0.1 -4,-2.7 1,-0.1 -1,-0.4 -0.430 59.1-125.0 -76.2 147.9 36.5 3.0 31.0 24 24 A R E -AB 18 612A 91 588,-0.6 588,-2.9 -6,-0.2 2,-0.3 -0.744 33.7-168.7 -89.9 140.2 36.4 -0.6 29.9 25 25 A S E -AB 17 611A 0 -8,-2.7 -8,-3.0 -2,-0.4 586,-0.2 -0.978 34.6-153.7-137.0 146.8 36.7 -1.1 26.2 26 26 A S E S+ 0 0 7 584,-2.6 -11,-2.5 -2,-0.3 2,-0.4 0.429 86.4 62.0 -91.4 -2.0 36.3 -3.9 23.6 27 27 A V E -A 14 0A 1 583,-0.4 2,-0.5 -13,-0.2 -13,-0.2 -0.970 61.1-161.5-131.0 143.2 38.7 -2.1 21.3 28 28 A Y E -A 13 0A 41 -15,-2.3 -15,-2.8 -2,-0.4 2,-0.4 -0.986 16.2-152.6-120.0 128.0 42.4 -1.0 21.5 29 29 A I E +A 12 0A 22 -2,-0.5 2,-0.2 -17,-0.2 -17,-0.2 -0.857 20.2 167.7-105.3 133.9 43.5 1.7 19.1 30 30 A D > + 0 0 9 -19,-2.5 3,-1.4 -2,-0.4 4,-0.2 -0.696 11.8 177.9-148.2 91.7 47.1 2.0 17.9 31 31 A X G > S+ 0 0 0 1,-0.3 3,-1.7 -2,-0.2 4,-0.4 0.677 75.6 78.7 -66.1 -18.4 47.8 4.3 15.0 32 32 A R G > S+ 0 0 1 1,-0.3 3,-1.6 2,-0.2 4,-0.4 0.811 79.3 70.0 -61.7 -28.4 51.5 3.5 15.3 33 33 A R G < S+ 0 0 71 -3,-1.4 3,-0.5 1,-0.3 -1,-0.3 0.678 86.0 66.7 -65.6 -14.9 50.8 0.2 13.5 34 34 A L G X S+ 0 0 0 -3,-1.7 3,-1.1 1,-0.2 7,-0.7 0.758 88.3 68.9 -75.4 -22.6 50.2 2.1 10.2 35 35 A L T < S+ 0 0 0 -3,-1.6 227,-2.7 -4,-0.4 -1,-0.2 0.780 85.0 67.6 -67.2 -27.5 53.8 3.1 10.1 36 36 A G T 3 S+ 0 0 0 237,-0.9 2,-0.3 -3,-0.5 -1,-0.3 0.255 98.8 55.2 -79.9 15.1 55.1 -0.3 9.4 37 37 A D <> - 0 0 53 -3,-1.1 4,-2.6 223,-0.1 3,-0.4 -0.883 66.9-153.4-149.6 112.9 53.6 -0.4 5.9 38 38 A E H > S+ 0 0 69 -2,-0.3 4,-2.2 1,-0.2 5,-0.2 0.878 99.2 56.4 -53.6 -40.0 54.2 2.2 3.3 39 39 A S H > S+ 0 0 71 1,-0.2 4,-1.3 2,-0.2 -1,-0.2 0.927 112.2 42.5 -58.2 -44.3 50.9 1.5 1.7 40 40 A S H > S+ 0 0 5 -3,-0.4 4,-2.0 2,-0.2 -2,-0.2 0.930 115.1 44.9 -71.1 -48.9 49.1 2.3 5.0 41 41 A Y H X S+ 0 0 0 -4,-2.6 4,-2.7 -7,-0.7 -1,-0.2 0.836 110.0 57.2 -69.0 -26.1 51.0 5.3 6.1 42 42 A S H X S+ 0 0 21 -4,-2.2 4,-2.3 -5,-0.3 -1,-0.2 0.894 106.6 48.3 -68.6 -38.6 50.8 6.9 2.7 43 43 A V H X S+ 0 0 48 -4,-1.3 4,-2.0 -5,-0.2 -2,-0.2 0.932 113.6 48.8 -64.3 -43.7 47.0 6.6 2.7 44 44 A A H X S+ 0 0 0 -4,-2.0 4,-2.2 1,-0.2 -2,-0.2 0.907 110.5 49.9 -62.1 -44.9 47.1 8.2 6.2 45 45 A L H X S+ 0 0 0 -4,-2.7 4,-2.4 1,-0.2 -1,-0.2 0.904 109.5 51.1 -63.0 -42.1 49.4 11.0 5.1 46 46 A D H X S+ 0 0 80 -4,-2.3 4,-2.1 1,-0.2 -1,-0.2 0.894 111.2 48.0 -63.4 -38.9 47.3 11.8 2.1 47 47 A L H X S+ 0 0 30 -4,-2.0 4,-2.6 2,-0.2 5,-0.3 0.871 110.7 51.5 -69.0 -36.6 44.1 12.0 4.3 48 48 A L H X S+ 0 0 0 -4,-2.2 4,-2.4 2,-0.2 -2,-0.2 0.900 110.1 48.8 -65.8 -40.7 46.0 14.2 6.8 49 49 A L H X S+ 0 0 38 -4,-2.4 4,-1.2 2,-0.2 -2,-0.2 0.915 111.9 51.1 -64.9 -40.8 47.1 16.6 4.0 50 50 A E H < S+ 0 0 147 -4,-2.1 3,-0.3 1,-0.2 -2,-0.2 0.938 116.2 37.2 -61.7 -51.8 43.5 16.7 2.8 51 51 A V H < S+ 0 0 39 -4,-2.6 -1,-0.2 1,-0.2 -2,-0.2 0.814 109.9 62.5 -73.3 -29.0 41.8 17.5 6.1 52 52 A G H >X S+ 0 0 4 -4,-2.4 4,-1.7 -5,-0.3 3,-0.7 0.809 85.6 166.1 -63.7 -28.5 44.7 19.8 7.1 53 53 A G H 3X - 0 0 27 -4,-1.2 4,-2.2 -3,-0.3 3,-0.2 -0.046 59.0 -21.4 50.6-144.6 43.7 21.9 4.1 54 54 A Q H 3> S+ 0 0 115 1,-0.2 4,-2.3 2,-0.2 -1,-0.2 0.785 131.7 65.2 -69.4 -28.4 45.0 25.4 3.6 55 55 A D H <> S+ 0 0 53 -3,-0.7 4,-0.6 2,-0.2 -1,-0.2 0.939 111.5 36.3 -59.0 -46.7 45.9 25.8 7.3 56 56 A L H >< S+ 0 0 1 -4,-1.7 3,-0.8 1,-0.2 -2,-0.2 0.930 115.8 55.0 -71.0 -44.9 48.5 23.1 7.0 57 57 A A H 3< S+ 0 0 30 -4,-2.2 -2,-0.2 1,-0.2 -1,-0.2 0.776 116.0 36.9 -60.7 -29.7 49.5 24.1 3.5 58 58 A R H 3< S+ 0 0 183 -4,-2.3 -1,-0.2 -5,-0.1 -2,-0.2 0.481 88.5 123.7-102.5 -3.8 50.3 27.7 4.5 59 59 A S << - 0 0 1 -3,-0.8 50,-0.2 -4,-0.6 3,-0.1 -0.208 61.8-138.9 -57.3 144.9 51.7 26.9 8.0 60 60 A S S S- 0 0 44 48,-3.3 2,-0.3 1,-0.2 49,-0.2 0.720 85.8 -0.2 -77.6 -19.0 55.2 28.2 8.7 61 61 A A E -c 109 0B 0 47,-0.8 49,-1.4 21,-0.1 2,-0.5 -0.984 65.6-124.6-164.0 155.6 55.9 24.9 10.4 62 62 A V E -cd 110 84B 0 21,-2.3 23,-1.8 -2,-0.3 2,-0.5 -0.926 28.1-159.1-106.0 129.7 54.4 21.5 11.3 63 63 A I E -cd 111 85B 0 47,-2.8 49,-2.9 -2,-0.5 2,-0.3 -0.945 2.0-153.3-114.9 127.1 54.6 20.7 15.0 64 64 A G E -cd 112 86B 0 21,-2.5 23,-2.1 -2,-0.5 2,-1.1 -0.719 16.5-130.5 -98.9 147.1 54.3 17.2 16.3 65 65 A V E >> - d 0 87B 6 47,-2.4 4,-2.1 -2,-0.3 5,-0.5 -0.843 55.7 -94.4 -90.6 93.9 53.1 16.2 19.8 66 66 A A T 45S- 0 0 3 21,-2.2 2,-0.2 -2,-1.1 20,-0.0 -0.435 86.3 -1.3 -49.0 126.2 55.8 13.8 20.8 67 67 A T T >45S+ 0 0 36 -2,-0.3 3,-1.1 21,-0.1 -1,-0.1 -0.943 128.9 38.4-124.2 -48.4 55.9 10.8 20.5 68 68 A G T 345S+ 0 0 15 301,-0.3 4,-0.4 1,-0.3 -2,-0.2 0.746 126.3 36.8 -29.8 -57.3 52.8 9.5 19.0 69 69 A G T 3X5S+ 0 0 0 -4,-2.1 4,-2.5 1,-0.2 -1,-0.3 0.534 95.6 89.0 -89.4 -3.4 52.4 12.3 16.6 70 70 A L H <> S+ 0 0 0 0, 0.0 4,-2.5 0, 0.0 -1,-0.2 0.927 115.3 48.9 -60.7 -39.9 56.5 10.4 12.9 72 72 A W H > S+ 0 0 0 -4,-0.4 4,-2.3 2,-0.2 -2,-0.2 0.921 112.2 46.9 -63.3 -45.4 53.4 11.9 11.3 73 73 A A H X S+ 0 0 0 -4,-2.5 4,-2.6 1,-0.2 -1,-0.2 0.912 112.9 49.9 -63.4 -43.4 54.5 15.5 11.8 74 74 A A H X S+ 0 0 0 -4,-2.8 4,-2.0 1,-0.2 -2,-0.2 0.909 112.1 47.1 -62.8 -42.9 58.0 14.7 10.4 75 75 A X H X S+ 0 0 0 -4,-2.5 4,-2.1 -5,-0.2 -1,-0.2 0.878 113.0 48.9 -67.5 -37.8 56.5 13.0 7.3 76 76 A L H X S+ 0 0 0 -4,-2.3 4,-2.1 2,-0.2 -2,-0.2 0.898 109.7 52.2 -67.0 -41.3 54.1 15.8 6.7 77 77 A A H X>S+ 0 0 0 -4,-2.6 5,-2.8 1,-0.2 4,-0.5 0.911 111.1 47.0 -62.0 -42.1 56.9 18.4 7.0 78 78 A L H ><5S+ 0 0 28 -4,-2.0 3,-0.8 3,-0.2 -1,-0.2 0.918 111.0 51.4 -66.0 -42.9 59.0 16.5 4.5 79 79 A R H 3<5S+ 0 0 106 -4,-2.1 -1,-0.2 1,-0.2 -2,-0.2 0.870 117.9 39.1 -61.5 -35.9 56.1 16.2 2.1 80 80 A L H 3<5S- 0 0 50 -4,-2.1 -1,-0.2 -5,-0.1 -2,-0.2 0.417 106.3-125.7 -94.9 1.4 55.4 19.9 2.4 81 81 A S T <<5 + 0 0 74 -3,-0.8 -3,-0.2 -4,-0.5 135,-0.2 0.901 65.3 140.8 53.8 42.1 59.1 20.9 2.4 82 82 A K < - 0 0 36 -5,-2.8 -1,-0.2 -6,-0.2 -21,-0.1 -0.774 59.2 -90.7-116.2 159.6 58.4 22.8 5.7 83 83 A P - 0 0 25 0, 0.0 -21,-2.3 0, 0.0 2,-0.4 -0.288 40.8-144.1 -63.3 149.5 60.2 23.3 9.0 84 84 A L E +d 62 0B 0 129,-2.9 2,-0.3 -23,-0.2 -21,-0.2 -0.911 24.1 164.2-119.6 147.6 59.5 20.7 11.7 85 85 A G E -d 63 0B 0 -23,-1.8 -21,-2.5 -2,-0.4 2,-0.3 -0.938 10.2-170.1-149.6 166.1 59.2 21.2 15.4 86 86 A Y E -dE 64 101B 0 15,-1.6 15,-2.1 -2,-0.3 2,-0.4 -0.987 15.7-143.0-162.5 155.5 57.8 19.1 18.3 87 87 A V E -d 65 0B 8 -23,-2.1 -21,-2.2 -2,-0.3 13,-0.1 -0.986 16.4-160.6-124.8 130.7 56.9 19.2 22.0 88 88 A R - 0 0 79 -2,-0.4 2,-0.7 11,-0.4 11,-0.2 -0.850 27.5-111.6-112.1 146.6 57.4 16.3 24.3 89 89 A P - 0 0 46 0, 0.0 2,-0.2 0, 0.0 3,-0.0 -0.664 41.1-150.7 -76.6 112.7 55.8 15.7 27.7 90 90 A E - 0 0 54 -2,-0.7 2,-0.2 1,-0.1 280,-0.1 -0.559 21.6 -99.5 -87.1 149.3 58.6 16.1 30.2 91 91 A R > - 0 0 178 -2,-0.2 3,-0.6 1,-0.1 5,-0.3 -0.468 50.0-119.7 -64.6 134.4 58.9 14.4 33.6 92 92 A K T 3 S+ 0 0 83 393,-0.6 -1,-0.1 1,-0.2 395,-0.0 -0.453 80.9 28.7 -85.5 150.6 57.7 16.8 36.3 93 93 A G T 3 S+ 0 0 63 -2,-0.2 -1,-0.2 3,-0.1 -2,-0.0 0.569 85.7 102.4 85.0 11.7 59.6 18.3 39.3 94 94 A H S < S- 0 0 138 -3,-0.6 -2,-0.1 2,-0.1 -3,-0.0 0.409 96.6-104.4-106.0 1.1 63.1 18.1 37.8 95 95 A G + 0 0 73 1,-0.3 2,-0.2 0, 0.0 -3,-0.1 0.653 65.3 158.7 85.3 16.1 63.6 21.7 36.9 96 96 A T - 0 0 73 -5,-0.3 2,-0.5 1,-0.0 -1,-0.3 -0.513 31.6-147.6 -75.9 140.2 63.1 21.1 33.2 97 97 A L - 0 0 148 -2,-0.2 -1,-0.0 1,-0.0 0, 0.0 -0.940 21.0-121.1-110.1 127.1 62.1 24.0 31.0 98 98 A S + 0 0 59 -2,-0.5 2,-0.3 -8,-0.1 -11,-0.0 -0.179 35.6 175.2 -64.1 157.5 59.8 23.2 28.0 99 99 A Q - 0 0 76 -11,-0.2 2,-1.1 -13,-0.1 -11,-0.4 -0.988 40.4-109.8-159.9 158.4 60.9 24.0 24.5 100 100 A V > - 0 0 11 -2,-0.3 3,-1.9 28,-0.1 -13,-0.2 -0.822 43.9-140.3 -94.6 97.1 59.8 23.6 20.9 101 101 A E B 3 S+E 86 0B 2 -15,-2.1 -15,-1.6 -2,-1.1 110,-0.0 -0.323 85.9 34.4 -60.3 137.5 62.4 21.1 19.8 102 102 A G T 3 S- 0 0 2 -17,-0.2 112,-0.9 1,-0.2 -1,-0.2 -0.020 97.6-130.3 104.9 -28.4 63.7 21.8 16.3 103 103 A D < - 0 0 43 -3,-1.9 -1,-0.2 -17,-0.2 -2,-0.1 0.902 31.9-169.7 48.0 53.9 63.4 25.5 16.9 104 104 A P - 0 0 0 0, 0.0 111,-0.1 0, 0.0 -1,-0.1 -0.334 36.9 -78.9 -68.2 158.1 61.5 26.3 13.7 105 105 A P - 0 0 68 0, 0.0 -45,-0.0 0, 0.0 -2,-0.0 -0.228 56.6 -92.4 -57.6 145.3 61.1 30.1 12.8 106 106 A K S S+ 0 0 120 -3,-0.1 2,-0.3 28,-0.1 28,-0.2 -0.319 80.2 93.1 -58.2 138.3 58.4 32.0 14.8 107 107 A G S S- 0 0 23 26,-0.2 28,-2.0 -3,-0.1 2,-0.4 -0.978 78.4 -33.9 164.6-167.3 55.1 32.0 12.9 108 108 A R E + f 0 135B 118 -2,-0.3 -48,-3.3 26,-0.2 -47,-0.8 -0.676 60.9 178.2 -84.7 130.4 51.8 30.3 12.5 109 109 A V E -cf 61 136B 1 26,-2.9 28,-0.5 -2,-0.4 29,-0.5 -0.820 26.3-135.7-128.3 168.0 52.0 26.5 12.9 110 110 A V E -cf 62 138B 2 -49,-1.4 -47,-2.8 -2,-0.3 2,-0.3 -0.989 22.0-139.5-122.9 131.3 49.7 23.5 13.0 111 111 A V E -cf 63 139B 1 27,-1.9 29,-3.0 -2,-0.4 2,-0.4 -0.720 16.7-159.2 -93.1 145.1 50.0 20.8 15.7 112 112 A V E +cf 64 140B 2 -49,-2.9 -47,-2.4 -2,-0.3 2,-0.3 -0.980 17.3 163.5-129.2 134.3 49.8 17.2 14.8 113 113 A D E - 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