==== Secondary Structure Definition by the program DSSP, updated CMBI version by ElmK / April 1,2000 ==== DATE=20-JUL-2011 . REFERENCE W. KABSCH AND C.SANDER, BIOPOLYMERS 22 (1983) 2577-2637 . HEADER OXIDOREDUCTASE 17-DEC-08 2ZWN . COMPND 2 MOLECULE: TWO-DOMAIN TYPE LACCASE; . SOURCE 2 ORGANISM_SCIENTIFIC: METAGENOMES; . AUTHOR H.KOMORI,K.MIYAZAKI,Y.HIGUCHI . 949 3 0 0 0 TOTAL NUMBER OF RESIDUES, NUMBER OF CHAINS, NUMBER OF SS-BRIDGES(TOTAL,INTRACHAIN,INTERCHAIN) . 29845.0 ACCESSIBLE SURFACE OF PROTEIN (ANGSTROM**2) . 664 70.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(J) , SAME NUMBER PER 100 RESIDUES . 87 9.2 TOTAL NUMBER OF HYDROGEN BONDS IN PARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 258 27.2 TOTAL NUMBER OF HYDROGEN BONDS IN ANTIPARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-5), SAME NUMBER PER 100 RESIDUES . 3 0.3 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-4), SAME NUMBER PER 100 RESIDUES . 18 1.9 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-3), SAME NUMBER PER 100 RESIDUES . 2 0.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-2), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-1), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+0), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+1), SAME NUMBER PER 100 RESIDUES . 132 13.9 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+2), SAME NUMBER PER 100 RESIDUES . 114 12.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+3), SAME NUMBER PER 100 RESIDUES . 23 2.4 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+4), SAME NUMBER PER 100 RESIDUES . 6 0.6 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+5), SAME NUMBER PER 100 RESIDUES . 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 *** HISTOGRAMS OF *** . 0 0 0 5 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 RESIDUES PER ALPHA HELIX . 3 0 3 3 0 0 3 0 3 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PARALLEL BRIDGES PER LADDER . 18 0 9 9 6 3 6 3 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 ANTIPARALLEL BRIDGES PER LADDER . 0 0 0 6 0 6 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 LADDERS PER SHEET . # RESIDUE AA STRUCTURE BP1 BP2 ACC N-H-->O O-->H-N N-H-->O O-->H-N TCO KAPPA ALPHA PHI PSI X-CA Y-CA Z-CA 1 2 A A 0 0 73 0, 0.0 42,-2.0 0, 0.0 2,-0.5 0.000 360.0 360.0 360.0 148.0 82.9 64.5 41.8 2 3 A E E -a 43 0A 138 40,-0.2 2,-0.5 42,-0.1 42,-0.2 -0.745 360.0-173.8 -82.5 123.3 84.0 67.5 43.9 3 4 A R E -a 44 0A 67 40,-3.1 42,-2.5 -2,-0.5 2,-0.4 -0.960 1.6-171.1-125.6 112.6 84.0 70.6 41.7 4 5 A E E -a 45 0A 104 -2,-0.5 2,-0.3 40,-0.2 42,-0.2 -0.893 4.9-178.2-111.2 132.6 85.3 73.8 43.2 5 6 A F E -a 46 0A 23 40,-3.1 42,-2.8 -2,-0.4 2,-0.5 -0.944 18.9-145.3-129.7 147.1 85.0 77.2 41.5 6 7 A D E +a 47 0A 118 -2,-0.3 2,-0.3 40,-0.2 42,-0.2 -0.967 25.4 176.2-112.5 124.7 86.1 80.7 42.4 7 8 A M E -a 48 0A 1 40,-3.0 42,-2.1 -2,-0.5 2,-0.3 -0.924 11.9-160.1-126.4 152.1 83.9 83.6 41.3 8 9 A T E -a 49 0A 38 -2,-0.3 20,-2.7 40,-0.2 2,-0.5 -0.840 15.6-134.9-121.4 165.1 83.9 87.3 41.8 9 10 A I E -aB 50 27A 0 40,-1.9 42,-1.9 -2,-0.3 2,-0.3 -0.997 33.2-174.8-118.6 125.4 81.3 90.1 41.6 10 11 A E E -aB 51 26A 31 16,-2.2 16,-1.8 -2,-0.5 2,-0.8 -0.900 30.9-124.0-126.4 148.7 82.6 93.1 39.6 11 12 A E E + B 0 25A 55 40,-0.5 306,-0.5 -2,-0.3 2,-0.3 -0.839 52.3 148.6 -88.2 111.0 81.4 96.6 38.8 12 13 A V E - B 0 24A 1 12,-1.7 12,-3.0 -2,-0.8 2,-0.4 -0.908 43.4-124.3-137.6 166.4 81.6 96.9 35.0 13 14 A T E -CB 315 23A 29 302,-2.0 302,-3.0 -2,-0.3 2,-0.3 -0.963 25.6-168.3-114.4 131.1 79.8 98.6 32.1 14 15 A I E - B 0 22A 11 8,-2.6 8,-3.0 -2,-0.4 2,-0.8 -0.851 24.9-122.7-116.7 152.7 78.3 96.6 29.2 15 16 A K E + B 0 21A 101 -2,-0.3 6,-0.2 6,-0.2 -2,-0.0 -0.865 33.2 169.7 -94.1 108.7 77.0 97.6 25.8 16 17 A V E - 0 0 0 4,-2.2 157,-2.8 -2,-0.8 -1,-0.2 0.862 66.7 -7.1 -86.0 -41.8 73.4 96.4 25.7 17 18 A A E > S- B 0 20A 0 3,-1.4 3,-2.7 155,-0.2 -1,-0.3 -0.936 94.6 -67.7-145.9 164.8 72.3 98.2 22.5 18 19 A P T 3 S+ 0 0 68 0, 0.0 3,-0.1 0, 0.0 158,-0.0 -0.446 127.9 9.3 -55.0 121.7 73.7 100.8 20.1 19 20 A G T 3 S+ 0 0 77 -2,-0.2 2,-0.4 1,-0.1 -4,-0.0 0.539 109.6 101.4 79.6 7.1 73.8 104.0 22.3 20 21 A L E < +B 17 0A 9 -3,-2.7 -4,-2.2 2,-0.0 -3,-1.4 -0.858 41.1 172.6-126.6 98.8 73.0 102.1 25.5 21 22 A D E -B 15 0A 88 -2,-0.4 2,-0.4 -6,-0.2 -6,-0.2 -0.838 18.7-159.2 -93.2 141.9 75.9 101.4 27.9 22 23 A Y E -B 14 0A 0 -8,-3.0 -8,-2.6 -2,-0.4 2,-0.5 -0.985 17.2-131.3-133.9 128.8 74.7 99.9 31.2 23 24 A K E +B 13 0A 39 128,-2.8 2,-0.3 -2,-0.4 -10,-0.2 -0.663 41.2 169.6 -75.0 125.0 76.3 99.7 34.6 24 25 A V E -B 12 0A 0 -12,-3.0 -12,-1.7 -2,-0.5 2,-0.7 -0.823 44.2-122.9-132.3 167.4 76.0 96.2 36.0 25 26 A F E -B 11 0A 0 90,-2.5 2,-0.5 -2,-0.3 -14,-0.2 -0.874 43.3-165.4-109.1 91.2 77.3 93.9 38.7 26 27 A G E > -B 10 0A 0 -16,-1.8 3,-2.3 -2,-0.7 -16,-2.2 -0.681 23.1-127.0 -90.8 126.9 78.8 91.2 36.6 27 28 A F E > S-BD 9 30A 0 3,-0.6 3,-1.4 -2,-0.5 -18,-0.2 -0.556 105.7 -2.2 -62.6 125.0 79.7 87.8 38.0 28 29 A N T 3 S- 0 0 25 -20,-2.7 -1,-0.3 -2,-0.3 -19,-0.1 0.792 132.5 -67.8 51.2 31.7 83.4 87.5 36.8 29 30 A G T < S+ 0 0 3 -3,-2.3 285,-2.7 -21,-0.2 2,-0.3 0.771 112.4 101.3 65.2 27.9 82.8 90.9 35.1 30 31 A Q B < -DE 27 313A 0 -3,-1.4 -3,-0.6 283,-0.3 283,-0.2 -0.985 65.5-122.2-142.0 153.9 80.4 89.6 32.5 31 32 A V S S+ 0 0 2 281,-2.1 2,-0.2 -2,-0.3 88,-0.1 -0.987 102.9 39.1-135.5 123.5 76.6 89.4 31.8 32 33 A P S S- 0 0 0 0, 0.0 87,-0.2 0, 0.0 277,-0.2 0.616 108.3-122.9 -53.4 157.2 75.4 86.6 31.4 33 34 A G - 0 0 0 85,-2.8 -5,-0.1 -2,-0.2 -2,-0.1 -0.139 45.1 -62.4 -64.7 171.0 77.6 85.3 34.2 34 35 A P - 0 0 0 0, 0.0 2,-0.4 0, 0.0 274,-0.2 -0.212 55.7-107.1 -56.8 144.9 79.9 82.3 33.6 35 36 A L E -G 307 0B 0 272,-2.2 272,-2.6 84,-0.1 2,-0.5 -0.635 28.9-160.9 -68.9 125.6 78.4 79.0 32.6 36 37 A I E -Gh 306 121B 1 84,-2.2 86,-2.9 -2,-0.4 2,-0.5 -0.973 13.3-179.8-108.0 121.0 78.6 76.4 35.4 37 38 A H E +Gh 305 122B 9 268,-2.9 268,-2.7 -2,-0.5 2,-0.2 -0.981 11.8 151.8-128.5 115.0 78.3 73.0 33.9 38 39 A V E - h 0 123B 0 84,-2.6 86,-2.8 -2,-0.5 2,-0.3 -0.790 39.5-102.3-135.5 175.9 78.3 69.9 36.2 39 40 A Q E > - h 0 124B 67 -2,-0.2 3,-2.6 84,-0.2 55,-0.2 -0.815 46.2 -88.3-105.8 145.1 77.0 66.4 36.5 40 41 A E T 3 S+ 0 0 57 84,-2.9 55,-0.2 -2,-0.3 -1,-0.1 -0.235 116.3 23.2 -44.8 126.8 74.1 65.1 38.5 41 42 A G T 3 S+ 0 0 25 53,-2.4 -1,-0.3 1,-0.3 54,-0.2 0.265 88.0 140.3 95.0 -10.2 75.4 64.2 42.0 42 43 A D < - 0 0 3 -3,-2.6 52,-3.0 52,-0.4 2,-0.7 -0.299 54.2-126.0 -62.4 147.8 78.5 66.4 41.9 43 44 A D E -aF 2 93A 64 -42,-2.0 -40,-3.1 50,-0.2 2,-0.4 -0.897 31.0-157.6 -95.4 114.8 79.5 68.2 45.1 44 45 A V E -aF 3 92A 1 48,-3.2 48,-2.1 -2,-0.7 2,-0.4 -0.790 14.7-175.5-103.0 134.1 79.8 71.9 44.1 45 46 A I E -aF 4 91A 11 -42,-2.5 -40,-3.1 -2,-0.4 2,-0.5 -0.997 2.8-175.0-125.5 124.2 81.8 74.5 46.0 46 47 A V E -aF 5 90A 0 44,-2.7 44,-2.9 -2,-0.4 2,-0.7 -0.967 9.4-161.2-124.0 116.2 81.7 78.1 44.9 47 48 A N E -aF 6 89A 30 -42,-2.8 -40,-3.0 -2,-0.5 2,-0.5 -0.872 16.3-163.7 -95.1 116.8 83.9 80.7 46.6 48 49 A V E -aF 7 88A 1 40,-3.2 40,-2.0 -2,-0.7 2,-0.5 -0.897 10.4-168.4-110.2 126.7 82.5 84.1 46.0 49 50 A T E -aF 8 87A 45 -42,-2.1 -40,-1.9 -2,-0.5 2,-1.0 -0.969 16.6-142.0-113.4 126.5 84.4 87.4 46.4 50 51 A N E +a 9 0A 0 36,-2.8 35,-2.3 -2,-0.5 -40,-0.1 -0.753 24.9 171.0 -90.2 99.6 82.5 90.7 46.3 51 52 A N E +a 10 0A 80 -42,-1.9 -40,-0.5 -2,-1.0 2,-0.2 -0.027 51.6 91.4 -96.8 30.5 84.6 93.3 44.5 52 53 A T S S- 0 0 10 2,-0.3 33,-0.4 -42,-0.2 4,-0.1 -0.637 85.2-112.7-115.4 177.8 81.7 95.8 44.4 53 54 A S S S+ 0 0 75 -2,-0.2 31,-0.2 31,-0.1 -2,-0.0 0.433 93.4 61.6 -88.2 -2.7 80.6 98.7 46.6 54 55 A L S S- 0 0 49 29,-0.1 -2,-0.3 28,-0.0 31,-0.2 -0.868 87.0 -98.3-129.5 155.8 77.3 96.9 47.6 55 56 A P + 0 0 31 0, 0.0 2,-0.3 0, 0.0 28,-0.2 -0.455 45.2 165.4 -73.3 145.8 76.1 93.7 49.3 56 57 A H B -K 82 0C 0 26,-1.8 26,-3.0 -2,-0.1 2,-0.3 -0.897 18.6-163.9-143.0 172.0 75.1 90.7 47.3 57 58 A T - 0 0 0 -2,-0.3 2,-0.4 24,-0.2 49,-0.2 -0.877 21.9-122.8-138.8 178.2 74.4 87.0 47.6 58 59 A I - 0 0 0 -2,-0.3 2,-0.4 46,-0.2 46,-0.2 -0.965 20.4-167.9-124.7 119.5 74.1 84.1 45.2 59 60 A H E -I 103 0B 1 44,-3.4 44,-2.4 -2,-0.4 2,-0.9 -0.871 16.6-141.8 -96.3 138.2 70.9 82.0 45.1 60 61 A W E > -I 102 0B 0 12,-0.5 3,-1.2 -2,-0.4 2,-0.5 -0.851 20.5-147.1-104.8 96.3 71.1 78.8 43.1 61 62 A H E 3 S+I 101 0B 0 40,-1.9 40,-1.8 -2,-0.9 60,-0.1 -0.558 80.6 15.4 -79.1 117.2 67.7 78.7 41.5 62 63 A G T 3 S+ 0 0 7 -2,-0.5 2,-0.4 1,-0.4 -1,-0.3 0.028 92.8 121.1 108.3 -23.5 66.4 75.1 41.1 63 64 A V < - 0 0 4 -3,-1.2 461,-0.6 38,-0.3 -1,-0.4 -0.622 54.8-151.5 -76.0 125.4 69.0 73.6 43.5 64 65 A H - 0 0 42 -2,-0.4 3,-0.2 29,-0.2 -1,-0.1 0.793 15.2-144.6 -66.9 -31.1 67.1 71.9 46.4 65 66 A Q > - 0 0 0 28,-0.3 3,-2.2 1,-0.2 2,-0.5 0.931 19.4-158.5 59.1 49.0 69.9 72.5 48.9 66 67 A K T 3 S- 0 0 109 1,-0.3 -1,-0.2 27,-0.3 27,-0.2 -0.451 76.4 -12.9 -63.0 108.1 69.0 69.1 50.5 67 68 A G T 3 S+ 0 0 65 -2,-0.5 -1,-0.3 496,-0.2 -2,-0.1 0.603 128.4 83.1 71.2 13.9 70.5 69.4 54.0 68 69 A T X + 0 0 45 -3,-2.2 3,-2.6 -4,-0.1 -2,-0.2 -0.107 41.1 141.0-137.9 36.3 72.4 72.6 52.9 69 70 A W G > + 0 0 29 494,-0.4 3,-2.0 1,-0.3 4,-0.2 0.747 64.9 71.3 -56.3 -26.2 69.7 75.3 53.3 70 71 A R G 3 S+ 0 0 105 1,-0.3 -1,-0.3 2,-0.1 3,-0.2 0.563 95.3 54.7 -70.4 -6.1 72.3 77.7 54.7 71 72 A S G < S+ 0 0 16 -3,-2.6 -1,-0.3 1,-0.1 -11,-0.2 0.197 75.5 103.2-102.6 12.2 73.8 77.9 51.1 72 73 A D S < S- 0 0 0 -3,-2.0 -12,-0.5 -13,-0.1 -2,-0.1 0.723 83.1-132.5 -71.2 -21.3 70.5 78.9 49.6 73 74 A G + 0 0 0 1,-0.2 32,-0.2 -3,-0.2 -14,-0.1 0.759 54.0 140.8 81.1 29.3 71.7 82.5 49.3 74 75 A V >>> - 0 0 3 -16,-0.1 4,-2.9 -17,-0.1 3,-1.6 -0.912 48.6-130.1-108.4 118.1 68.8 84.5 50.7 75 76 A P B 345S+l 79 0D 36 0, 0.0 5,-0.2 0, 0.0 527,-0.1 -0.393 90.6 11.3 -70.4 139.1 69.6 87.4 52.9 76 77 A G T 345S+ 0 0 20 3,-2.1 526,-2.6 1,-0.1 527,-0.3 0.440 131.1 55.3 73.8 0.7 67.9 87.6 56.2 77 78 A V T <45S+ 0 0 6 -3,-1.6 521,-0.2 2,-0.5 -1,-0.1 0.636 126.1 3.8-124.3 -52.3 66.6 84.0 55.7 78 79 A T T <5S- 0 0 1 -4,-2.9 2,-0.3 1,-0.3 -6,-0.1 0.571 130.2 -5.2-115.3 -16.5 69.6 81.7 55.1 79 80 A Q B < -l 75 0D 4 -5,-1.6 -3,-2.1 521,-0.1 -2,-0.5 -0.979 69.6 -98.6-160.4 168.5 72.6 84.1 55.5 80 81 A Q - 0 0 119 -2,-0.3 -7,-0.0 -5,-0.2 3,-0.0 -0.683 61.5 -87.2 -84.8 152.1 73.6 87.6 56.1 81 82 A P - 0 0 42 0, 0.0 2,-1.0 0, 0.0 -24,-0.2 -0.115 36.5-106.9 -59.8 159.3 74.6 89.4 52.8 82 83 A I B -K 56 0C 4 -26,-3.0 -26,-1.8 -3,-0.1 3,-0.1 -0.804 46.0-143.7 -84.5 104.4 78.1 89.3 51.4 83 84 A E > - 0 0 91 -2,-1.0 3,-2.3 -28,-0.2 -33,-0.3 -0.201 33.4 -68.2 -71.1 163.3 79.3 92.8 52.2 84 85 A A T 3 S+ 0 0 44 1,-0.3 -1,-0.2 -31,-0.2 -33,-0.2 -0.262 122.7 9.4 -50.3 127.7 81.5 94.9 49.9 85 86 A G T 3 S+ 0 0 51 -35,-2.3 -1,-0.3 -33,-0.4 2,-0.1 0.310 105.8 115.2 84.7 -7.4 85.0 93.4 49.8 86 87 A D < - 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